Hoffeinsodes bifida, Wagner, Rüdiger, 2017

Wagner, Rüdiger, 2017, Synopsis of extinct Bruchomyiinae (Diptera, Psychodidae) from Burmese, Baltic and Dominican amber, with descriptions of new genera and species, Zootaxa 4320 (1), pp. 100-120 : 103-104

Hoffeinsodes bifida sp. nov.

( Figs. 2a–b )

Etymology. Bifida (lat.) = forked; describes the shape of posterolateral lobes of epandrium.

Diagnosis. Hoffeinsodes species with cross sclerite with 2 lateral tips, three dimensional glabrous part of gonostyli hardly longer than setose part, posterolateral lobes of epandrium sharp, bifid.

Material. Holotype male, no. 1515- 1 in coll. Hoffeins.

Geological stratum. Baltic amber.

State of preservation. Specimen with head, thorax and abdomen well conserved; palpi and antennae complete; some milky coating on the right side of thorax, more on the left side, and left antenna. Front legs, left middle leg, and right hind leg well preserved, remaining missing.

Syninclusions. Collembola.

Description. Head with oval eyes, distance between eyes approx. 8–10 facet diameters. Antenna with scape, pedicel and 14 flagellomeres; relative length of scape, pedicel and flagellomeres: 10-12-49 -34-29-27-29-25-23-21- 23-18-17-16-16-18; absolute length: 0.06-0.07-0.29-0.20-0.17-0.16-0.17-0.15-0.14-0.12-0.14-0.11-0.10-0.09- 0.09-0.11 mm; apical flagellomere with short apiculus. Palpus 4-segmented, basal segments partly hidden by milky coating, length not clearly discernable: approximate relative length of palp segments: 12-26-43 -67; approximate absolute length: 0.07-0.15- 0.25-0.39 mm.

Head length 0.48 mm. Length of thorax, abdomen, and genitalia 2.39 mm; thorax with swelling above halters with basal rings of setae.

Wing length 3.96 mm. Sc runs close to R and ends free in wing, with short sc-r cross vein close to tip; basally a humeral vein. First basal cell narrow, extends to middle of wing. R2+3 originates at about middle of first basal cell, R2 slightly longer than R2+3. Veins R4 and R5 originate nearby at front edge, M1 and M2 at hind edge of basal cell, no M1+2 vein; M3 and M4 originate at close quarters from middle of second basal cell, CuA and CuP run parallel, distally curved to hind wing margin.

Abdomen with 8 segments and genitalia; segments 1 and 6 narrower than segments 2– 5. Genitalia inverted by torsion of segments 7 and 8; the cross sclerite without posterolateral tips. Gonostylus bilobed, consisting of a morphologically ventral setose rounded and a short three-sided dorsal glabrous lobe with sharp tip; behind setose part a tip of an elongate rectangular sclerite (probably paramere), tip of its counterpart visible on the other side of genitalia. Aedeagus not discernable. Epandrium deep cup-shaped, ventral margin with lateral, apically bilobed prolongations. Distal edge of epandrium deeply arched; proctiger thin slightly sclerotized elongate with blunt tip, cerci small oval setose.

Comments. A Hoffeinsodes species distinguished from its congeners by the cross sclerite without distal tips, shape of the gonostylus, and apically cleft posterolateral lobes. Furcation of epandrial lobes is covered with milky coating. The closest relative of H. bifida is H. obtusa , both with capsule-like genitalia; in particular the cup-shaped epandrium makes the abdomen appear rounded, as though the apex of the abdomen is enclosed.

Figures

FIGURE 2. Hoffeinsodes bifida sp. n. a. wing; b. genitalia left lateral view; Hoffeinsodes cubicula sp. n.: c. genitalia right lateral view; d. dorsal view with cross sclerite, gonostyli and parameres. Abbreviations: c—cercus; ep—epandrium; fhg—fused hypandrium and gonocoxites; gst—gonostylus; pm—paramere; p + c—proctiger with cerci.