Cyana Walker, 1854

Bayarsaikhan, Ulziijargal & Bae, Yang-Seop, 2016, A review of the genus Cyana Walker, 1854 (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with description of new species, Zootaxa 4114 (4), pp. 447-463 : 448

Genus Cyana Walker, 1854

Cyana Walker, 1854 , List. Spec. Lepid. Insects Colln. Br. Mus. 2: 528. TS: Cyana detrita Walker, 1854 .

Doliche Walker, 1854, List. Spec. Lepid. Insects Colln. Br. Mus. 2: 529. TS: Doliche gelida Walker, 1854 . Isine Walker, 1854, List. Spec. Lepid. Insects Colln. Br. Mus. 2: 545. TS: Isine trigutta Walker, 1854 .

Bizone Walker, 1854 , List. Spec. Lepid. Insects Colln. Br. Mus. 2: 548. TS: Bizone perornata Walker, 1854 . Chionaema Herrich-Schäffer , [1855], Syst. Bearb. Schmett. Europ. 6 (68): 100. TS: Phalaena puella Drury, 1773 . Clerckia Aurivillius, 1882, K. svenska Vetensk Akad. Handl. (n.s.) 19 (5): 157. TS: Phalaena fulvia Linnaeus, 1758 . Exotrocha Meyrick, 1886, Proc. Linn. Soc. N.S.W. (2) 1 (3): 691. TS: Phalaena liboria Stoll , [1781].

Neobrocha Meyrick, 1886 , Proc. Linn. Soc. N.S.W. (2) 1 (3): 692. TS: Neobrocha phaeocyma Meyrick, 1886 . Sphragidium Butler, 1887, Ann. Mag. Nat. Hist. (5) 19 (111): 218. TS: Sphragidium miles Butler, 1887 .

Gnophrioides Heylaerts, 1891, C. R. Soc. Ent. Belge 35: 412. TS: Gnophrioides flaviplaga Heylaerts, 1891 . Leptothrix Heylaerts, 1892 , Ann. Soc. Ent. Belg. 36 (2): 47. TS: Leptothrix tettigonioides Heylaerts, 1891 . Macronola Kirby, 1892, Syn . Cat. Lepid. Het. 1: 299. (repl. Cyana Walker, 1854 ).

Adult. Proboscis well developed. Antennae ciliate in both sexes. Forewing with at least two red, brown or gray transverse fascia, a basal spot, plus at least one apical spot. A tuft of modified scales (secondary sexual features) between discal part of cell and costa in the males. Vein M2 of hindwings obsolescent ( Karisch, 2013).

Male genitalia. Uncus moderately long, broad; saccular process usually tapered and curved to an acute apex; aedeagus short, broad, and vesica ornamented by scobinated zones or clumps of spines ( Holloway, 2001).

Female genitalia. Appendix bursae large; corpus bursae wrinkled. Signum scobinated and often one or two in number, when present ( Holloway, 2001).

Remarks. Recently, African species of this genus were classified as Cyana by Karisch (2013), who suggested that the genus be subdivide into 24 subgenera, but this proposal is not followed herein.