Eurylychnus cylindricus Sloane, 1916

Lay, Stuart, 2023, A taxonomic review of the Australian genus Eurylychnus Bates (Coleoptera: Carabidae: Broscini), Zootaxa 5330 (2), pp. 247-264 : 252-253

Eurylychnus cylindricus Sloane, 1916

( Figs. 1B , 2B , 4D–F )

Eurylychnus cylindricus Sloane, 1916: 199 . Moore et al. 1987: 120 (catalogue). Häckel et al. 2010: 65 (checklist).

Type material: Lectotype, here designated: (1), Type, Barrington R., 5200 ft. Brush, T. G. S. 29.12.15 [handwritten] // Type // Eurylychnus cylindricus Sl. Id. By T. G. Sloane [handwritten and typed] // Eurylychnus cylindricus Sl. HOLOTYPE PJD [red label, handwritten and typed], ( ANIC) [Darlington placed a holotype label on the specimen but did not publish that action]. Paralectotypes, here designated: (2), Barrington R, 5200 ft. Brush, T. G. S., 29.12.15 [handwritten] ( ANIC); (1), Barrington R, 5200 ft. Brush, T. G. S., 29.12.15 [handwritten] // ♀ [handwritten symbol] ( ANIC); (1) [♀], Barrington R, 5200ft. Brush, T.G.S., 29.12.15 // Eurylychnus cylindricum Sl., Id. By T. G. Sloane // ANIC Specimen [green label] ( ANIC); (1), Barrington R, 5200ft Brush, T.G.S., 29.12.15 // Cotype 2569 [blue label] // Eurylychnus cylindricus Sl. Id. by T. G. Sloane // F. E. Wilson Collection ( MV); (1), Williams R 4500ft Brush, T.G.S., 27.12.15 // Pres [?] by T.G. Sloane 584[?] 12-6-16 // Eurylychnus cylindricus Sl, Id. by T. G. Sloane. // Cotype [blue label] // PARATYPE, T-11819, Eurylychnus cylindricus [blue label] ( MV); (1), Barrington R., 5200ft Brush, T. G. S., 29.12.15 // Pres by T. G. Sloane B09[?] // Eurylychnus cylindricus Sl. Id. by T. G. Sloane . // Cotype [blue label] // PARATYPE, T-11820, Eurylychnus cylindricus [blue label] ( MV); (1), Barrington Tops NSW, 29.12.1915, A. Musgrave, 4800ft // Eurylychnus cylindricus Sl Id. by A. M. Lea // Cotype [blue label] // PARATYPE, T-11821, Eurylychnus cylindricus [blue label] ( MV).

Other material examined (29): New South Wales: (2), Barrington Tops, H.J. Carter ( ANIC); (1) , Barrington Tops , v.1937, Few ( ANIC); (1) , Barrington Tops , 7.xi.1959, B.P. Moore ( ANIC); (6) [1 ♁, 5 unknown sex], Barrington Tops , 25–6.xi.1963, B.P. Moore ( ANIC); (2) , Barrington Tops , 24.iii.2001, C. Rojewski ( ANIC); (2) , Barrington Tops , i.1925, SU ZooExp ( ANIC); (1) , Barrington Tops NP, Devils Hole , 12.ii.2022, S. Lay ( SLC); (7) [4 ♁, 3 ♀], Barrington Tops NP, Honeysuckle area 12.ii.2022, S. Lay ( SLC); (1) [♀], Barrington Tops, Nothofagus forest , 8.ii.1996, S. Watkins ( ANIC); (3) [1 ♁, 2 ♀], Barrington Tops NP, Quarry Road 12.ii.2022, S. Lay ( SLC); (1) [♀], Barrington Tops NP, Wild Turkey Road , 12.ii.2022, S. Lay ( SLC); (1) , Eccleston , iii.1921, T.G. Sloane ( ANIC); (1) , Mount Royal Range, 36km E[ast] Moonan Flat , 7.xi.1982, J. Doyen ( ANIC) .

Diagnosis: Eurylychnus cylindricus has a large body size like E. regularis and E. blagravei , however E. cylindricus is narrower and more cylindrical in form ( Fig. 2B ), and its elytral striae and transverse impression behind the eyes are less distinct. Eurylychnus cylindricus is sympatric with E. ovipennis , however E. cylindricus has a larger body length (<16.5mm) and has only a single seta either side of the pronotum, unlike E. ovipennis which has 4 setae on either side of the pronotum.

Redescription: Body shiny black, cylindrical in form, legs black, sometimes reddish-black, palps piceous. Body length 16.5–20.4mm; maximum pronotum width 4.7–5.6mm; maximum elytra width 5.3–6.2mm.

Head: Vertex smooth, with sparse minute micropunctures. Transverse impression behind the eyes weak. Frontal impressions curving inwards anteriorly, joining the frontoclypeal suture and extending onto the clypeus where they terminate with a seta before the anterior margin. Clypeus anterior margin straight or weakly emarginated. Labrum anterior margin straight or weakly emarginated, with 6 evenly spaced setae on its dorsal anterior edge, with the outermost setae longer than the inner 4. Eyes with a single supraorbital seta. Mandibles with a seta in the scrobe. Median tooth of mentum bifid, relatively broad, with a basal seta each side; paramedian foveae present; lobes broad. Submentum with 2 setae. Stipes with 2 setae each, the inner seta more than double the length of the outer seta. Gula convex with some sparse wrinkles; gular sutures with small gular pits.

Thorax: Pronotum covered with sparse minute micropunctures, becoming reticulation of fine meshes on lateral margins; sometimes with transverse wrinkles from median impression; foveae present adjacent to the basal angles, sometimes joined by a transverse row of faint punctures; posterior margins gently rounded to base, lacking tubercles on basal angles; pronotum widest at or slightly above the midline, with 1 seta on the lateral margins at or just below the midline; median impression runs most of the length of the pronotum; pronotal carina borders the lateral margins, extending slightly onto the anterior margins and terminates posteriorly at the basal angles. Prosternum very convex, smooth or with some wrinkles, with a raised margin on anterior border. Protrochanters with 1 seta each. Males lacking squamose setae on ventral surface of protarsi ( Fig. 1B ). Metatrochanters without a seta. Mesocoxae with or without a seta. Metacoxae with 1 seta on anterior margin.

Elytra: Elytral disc convex and elongate; covered in reticulation of fine meshes, sometimes confined to the edges of each interstice in the apical half of the elytra; lateral elytral carina extending from humerus to apex. Parascutellar striole isolated from striae 1, although sometimes faint. Elytra with 8 striae; striae 1–7 weakly to moderately defined, stria 8 almost indistinct; striae 3 and 4 converging or sub-parallel basally and stria 5 begins at the humeral angle; interstices flat; striae sometimes with small obsolete punctures. Basal groove with 3 asetose obsolete punctures either side of the suture. Humerus simple.

Abdomen: Ventrites 3–5 lacking setae. Apical margin of ventrite 6 with a pair of paramedian setae.

Male genitalia: Aedeagus left paramere ( Fig. 4D ) broad, asetose, constricted dorsally in apical ¼ to form a long blunt denticle; right paramere ( Fig. 4E ) long, narrow, width uniform for most its length, only tapering to a point at the apex, setae present along the ventral margin of the apical half. Median lobe ( Fig. 4F ) weakly to moderately curved ventrally, slightly to moderately deflected to the right; apex blunt and rounded.

Figures

FIGURE 1. Ventral surface of male protarsi of A, Eurylychnus dyschirioides; B, Eurylychnus cylindricus.

FIGURE 2. Dorsal habitus of A, Eurylychnus blagravei; B, Eurylychnus cylindricus; C, Eurylychnus regularis; D, Eurylychnus victoriae. Scalebars are 2mm.

FIGURE 4. Male genitalia of Eurylychnus blagravei (A–C): A, left paramere, B, right paramere, C, median lobe left lateral; E. cylindricus (D–F): D, left paramere, E, right paramere, F, median lobe left lateral; E. regularis (G–I): G, left paramere, H, right paramere, I, median lobe left lateral; E. victoriae (J–L): J, left paramere, K, right paramere, L, median lobe left lateral. Scale bars are 0.5mm.