Manota obtecta, Hippa, Heikki, 2009

Hippa, Heikki, 2009, New species and new records of Manota Williston (Diptera, Mycetophilidae) from Thailand, Zootaxa 2017, pp. 1-33 : 13

Manota obtecta sp. n.

Figs 7 A, B, C

Male. Colour. Head dark brown, face and clypeus pale yellowish-brown, mouthparts pale yellowish, antenna dark brown, scapus, pedicellus and the two basalmost flagellomeres ventrally paler brown. Thorax brown, prothorax, anterior part of scutum and ventral part of preepisternum 2 paler. Legs pale yellowish, the apices of coxae 2 and 3 and their trochanters infuscated, femur 3 indistinctly infuscated on basal third. Wing unicolorous greyish-brown; haltere yellowish-brown with dark brown knob. Abdomen brown, sternites 1–4 paler yellowish. All the setosity yellowish or brownish, the thicker setae seeming darker than the finer setae and trichia. Head. Antennal flagellomere 4, Fig. 7 A. Palpomere 3 of maxillary palpus with apicomesial extension, with 4 apically expanded curved sensilla; palpomere 4 with parasegment. Number of strong postocular setae 10–12. Thorax. Anepisternum setose, with 22–38 setae, anterior basalare setose, with 10–17 setae, preepisternum 2 non-setose, laterotergite setose, with 20–26 setae, episternum 3 setose, with 7–12 setae. Wing. Length 1.6–1.8 mm. Hypopygium, Figs 7 B, C: Sternite 9 laterally fused with gonocoxa, extending posteriorly as far as the ventral posterior margin of gonocoxa, anterior margin with a shallow incision, the setae similar to ventral setae of gonocoxa. Parastylar lobe sickle-shaped, the apex directed obliquely posteriad, with several apical setae. Paraapodemal lobe absent. Dorsal mesial margin of gonocoxa sinuous, at middle with a plate-like lobe bearing 8–9 blunt-ended megasetae at its posterior margin, subapically with a large flattened, apically rounded megaseta. One juxtagonostylar seta present: an unmodified strong seta arising from a basal body which is about half the length of the seta. Gonostylus short and broad, with a subapical laterodorsal lobe, with rather short unmodified setae ventrally, largely non-setose dorsally, with two prominent long setae subapically at mesial margin, some of the setae on the lateral lobe conspicuously curved. Aedeagus with a narrow apical part and prominent lateral shoulders giving rise to basal part that is several times broader, the apex of aedeagus curved ventrad. Hypoproct large, posteriorly extending nearly as far as the apex of gonostylus, with ca. scattered 30 setae on each half. Cerci mesially separate.

Female unknown.

Discussion. M. obtecta is similar to M. clausa Hippa , known from the Malay Peninsula. It is distinguished e.g. by the following characters: 1) there is a large flat megaseta subapically at the mesial margin of gonocoxa; 2) the megasetae on the flat lobe at the middle of the dorsal mesial margin of the gonocoxa are larger and fewer in number; and 3) the parastylar lobe is flat and curved, not conical and straight. Even the conspicuously bilobed gonostylus probably distinguishes M. obtecta from M. clausa . I have restudied the types of the latter: the mounts of both the holotype and the paratype are such that the exact structure of the gonostylus is difficult to see, but it seems that there is only a slight indication of a subapical laterodorsal lobe.

Etymology. The name is from Latin, obtecta , covered over, and refers to the large sternite 9 which in ventral view covers part of the other mesial structures of the hypopygium.

Types. Holotype. Male, THAILAND, Nakhon Ratchasima, Khao Yai NP, Cobra zone near Fire protection office, 14o28.524’N 101o22.928’E, 757 m, Malaise trap, 12–19.vi.2007, Pong Sandao leg.T2224 (in QSBG). Paratypes. 1 male, same data as holotype except for 5–12.vi.2007, T2221 (in QSBG); 1 male, Phetchabun, Thung Salaeng Luang NP, Gang Wang Nam Yen, pine forest, 16o36.284’N 100o53.128’E, 749 m, Malaise trap 29.vi–6. vii.2007, Pongpitak & Sathit leg. T2066 (in SMNH).

Figures

FIGURE 7. Manota obtecta sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. Scale 0.10 mm.