Perissommatidae (Amorim, 1993)

Barahona-Segovia, Rodrigo M., Madriz, Isaí, González, Christian R. & Amorim, Dalton De Souza, 2024, An update on the knowledge and general understanding of the Chilean Diptera diversity, Zootaxa 5518 (1), pp. 1-87 : 11

Perissommatidae ( Fig. 13 )

This small family contains only five extant species of the genus Perissomma Colless ( Colless 1962, 1969). Only Perissomma congrua Colless has been reported for Chile and there are no more recent taxonomic studies on the family.

5. Suborder Bibionomorpha

According to Hennig (1973), two synapomorphies support the monophyly of the Bibionomorpha : conspicuous enlargement of the second latero-tergite, and an undivided postphragma of the thorax. Fitzgerald (2004) considered the presence of dorsal sclerite and ventrolateral apodemes as synapomorphies for this suborder. The family composition of Bibionomorpha remains controversial. Wood & Borkent (1989) disagreed with Hennig (1973) and removed the Scatopsoidea from the group based on larval features, and placed it in the Psychodomorpha . This was supported by Amorim (1994, 2000), and Amorim & Grimaldi (2006) based on thoracic pleural features. More recently, Bertone et al. (2008) and Wiegmann et al. (2011), based on molecular data, and Lambkin et al. (2013), based on morphological data, moved the Scatopsoidea back to the Bibionomorpha . There are also problems with the position of Axymyiidae placed as the sister group to Bibionomorpha s.str. or in a clade with Pachyneuridae and Perissommatidae ( Amorim 1993) or grouped with Culicomorpha ( Ševčík et al. 2016). Both these families have the plesiomorphic forked R 2+3, which is not forked in Anisopodidae and Cramptonomyiidae (with R 2+3 entirely lost in all remaining Bibionomorpha families).

Figures

FIGURES 8–13. Culicomorpha and Perissommatomorpha of Chile: (8) Forcipomyia sp. (Ceratopogonidae, Photo by Matías Gargiulo); (9) Chironomidae (photo by Matías Gargiulo); (10) Aedes (Ochlerotatus) albifasciatus (Macquart) (Culicidae, photo by Claudio Maureira); (11) Cnesia sp. (Simuliidae, photo by Claudio Maureira); (12) Gigantodax brophyi (Edwards) (Simuliidae, photo by Claudio Maureira), and (13) Perissoma cf. congrua (Perissommatidae, photo by Claudio Maureira).