Acilepis silhetensis (DC.) H.Rob., Proc. Biol. Soc. Washington 112(1): 227. 1999.

Bunwong, Sukhonthip, Chantaranothai, Pranom & Keeley, Sterling C., 2014, Revisions and key to the Vernonieae (Compositae) of Thailand, PhytoKeys 37, pp. 25-101 : 44-46

Acilepis silhetensis (DC.) H.Rob., Proc. Biol. Soc. Washington 112(1): 227. 1999.

Vernonia bracteata Wall., Numer. List [Wallich] no. 2921, comp. no. 31, nom. nud.

Decaneurum silhetense DC., Prodr. 5: 67. 1836.

Vernonia silhetensis (DC.) Hand.-Mazz., Symb. Sin. 7: 1084. 1936.

Type.

India, Silhet; Wallich 2921 (holotype: E!). Fig. 5H .

Description.

Perennial herbs, 1-3 m tall. Stems erect, inconspicuously ribbed, pilose. Leaves 7-12 by 2-4 cm, lanceolate or oblanceolate, margin serrate, apex acute or acuminate, base attenuate, subcoriaceous; both surfaces scabrous with whip-shaped hairs and capitate glands; lateral veins 5-10-paired; petioles up to 1 cm long. Capitulescences terminal, paniculate or solitary. Capitula campanulate, 15-20 mm long, pedunculate. Receptacle flat, 6-10 mm in diam., hairy. Involucres in 6-7 series, 11-18 mm long, 10-15 mm in diam., herbaceous, campanulate. Phyllaries dark purple or green with purple apex, margin piliferous, outer surface arachnoid lacking glands; the outer and the middle ones ovate, apex acuminate, upper half strongly reflexed; the inner ones ovate-lanceolate to oblong, apex acuminate or aristate. Florets 50-75; corollas funnelform, purple, glandular; corolla tubes 8-12 mm long; corolla lobes 3.5-5 mm long. Anthers 3.5-4.5 mm long, apical appendage acute, base obtuse. Styles purple, 10-12 mm long, branches 3.5-4.5 mm long. Achenes subterete, 4-5 mm long, 10-ribbed, glandular. Pappus bristles, the inner ones 6.5-8 mm long.

Distribution.

Thailand: Mae Hong Son, Chiang Mai, Lamphun, Lampang, Phetchabun, Loei, Khon Kaen. China, India, Bhutan, Myanmar, Laos, Myanmar.

Specimens examined.

Thailand, Chiang Mai, Queen Sirikit botanic garden, 18°53.89'N, 98°51.61'E, 10 Dec 2007, S. Bunwong 364 (KKU, US); Mae Hong Son, Pai, 20 Oct 1979, T. Shimizu, H. Toyokuni, H. Koyama, T. Yahama & T. Santisuk T-20113 (BKF, L); Chiang Mai, Doi Chiangdao, Nov 25 1963, Adisai 714 (BK); Doi Chiangdao, 24 Jul 1998, K. Buchuai 279 (AAU, BKF); Doi Chiangdao, 16 Dec 1983, N. Fukuoka & M. Ito T-35208 (BKF); Doi Chiangdao,3 Dec 1984, H. Koyama T-39781 (BKF, L); Doi Chiangdao, 5 Aug 1988, H. Koyama T- 61104 (BKF); Doi Chiangdao,29 Nov 1984, H. Koyama, T. Yahara, H. Nagamasu, W. Nanakorn & N. Nantasan T-39710 (AAU, BKF, L); Doi Chiangdao, 30 Nov 1984, H. Koyama, T. Yahara, H. Nagamasu, W. Nanakorn & N. Nantasan T- 39736 (BKF, L); Doi Chiangdao, 4 Nov 1995, J.F. Maxwell 95-1065 (CMU, BKF); Doi Chiangdao, 13 Oct 1931, Put 311 (BK, BM, K); Chiang Mai, Doi Sutep, 6 Oct 1982, Konta Th. Wongprasert & B. Sangkhachand 29741 (BKF); Doi Sutep,28 Nov 1984, H. Koyama, S. Mitsuta, T. Yahara & H. Nagamasu T-39671 (BKF, L); Doi Inthanon, 6 Dec 1984, S. Mitsuta, T. Yamada & H. Nagamazu T-46454 (BKF); Doi Inthanon, 1 Oct 1971, G. Murata, K. Iwatsuki, C. Phengklai & C. Charoenpol T-15497 (BKF); Doi Inthanon, 1 Oct 1971, G. Murata, K. Iwatsuki, C. Phengklai & C. Charoenpol T-15498 (BKF, P); Mae Rim, 21 Dec, 1985, Y. Paisooksantivatana 1645b-85 (BK); Mae Rim, 26 Dec 1987, R. Pooma 43 (BKF); Lampang, Jae son national park, 2 Dec 1995, J.F. Maxwell 95-1227 (BKF, L); Doi Khun Tan, 2 Jan 1985, H. Koyama & C. Phengklai T-39197 (BKF, L); Phetchabun, Namnao national park, 18 Nov 1973, J.F. Maxwell 73-636 (AAU, BK, BKF); Loei, Phu Kradung, 24 Dec 1991, Dee 195 (E); Phu Kradung, 21 Oct 1989, Din 125 (BKF); Phu Kradung, 19 Dec 1982, H. Koyama, H. Terao & Th. Wongprasert T-31330 (C, BKF); Phu Kradung, 5 Sep 1988, R. Pooma 62 (BKF); Phu Kradung, 21 Oct 1967, Prayad 1076 (BK); Khon Kaen, route to Nam Nao national park, 18 Nov 1979, T. Shimizu, H. Toyokuni, H. Koyama, T. Yahama & T. Santisuk T-22531 (BKF, L).

Diagnostic characters.

This species is distinguished by its strongly reflexed phyllaries and glandular achenes lacking trichomes.

Ecology.

Dipterocarp or pine-oak forest, alt. 700-1350 m; flowering August to January.

Vernacular names.

Phak Phet Kao Kum (ผักเผ็ดข้าวกํ่า), Ya Klung (หญ้าคลัง), Ya Hang Nok Kiew (หญ้าหางนกเขียว), Hudsakuen (หัสคึน).

Figures

Figure 5. Morphology of Vernonieae in Thailand 1. A Acilepis attenuata B Acilepis divergens C Acilepis namnaoensis D Acilepis ngaoensis E Acilepis peguensis F Acilepis principis G Acilepis saligna H Acilepis silhetensis I Acilepis squarrosa.