Pinodytes, Horn, 1880

Peck, Stewart B. & Newton, Alfred F., 2017, An Annotated Catalog of the Leiodidae (Coleoptera) of the Nearctic Region (Continental North America North of Mexico), The Coleopterists Bulletin 71 (2), pp. 211-258 : 215-217

Genus PINODYTES Horn, 1880

Pinodytes Horn, 1880: 248 . Type species: Catops cryptophagoides Mannerheim, 1852 (by monotypy). Portevin 1922: 2 (as synonym of Catopocerus ); Peck and Cook 2011: 14 (resurrected as valid genus).

Biology. All species are eyeless and flightless soil inhabitants (edaphobites), probably feed on subterranean fungi, and are sometimes collected in caves.

Distribution. W Nearctic (unglaciated lands west of the Mississippi River) (42 species). = Homeosoma Austin, 1880: 16 (attributed to Horn;

preoccupied, not Curtis, 1833). Type species:

Catops cryptophagoides Mannerheim, 1852 (by

monotypy). Hatch 1928: 72 (synonymy).

Note. As Homaeosoma in Neave (1939–40) = Typhloleiodes Hatch, 1935: 116 . Type species:

Typhloleiodes subterraneus Hatch, 1935 (by origi-

nal designation). Hatch 1957: 19 (synonymy).

P. angulatus Peck and Cook, 2011: 43 . Holotype male in CMNC. Type locality: Green Creek , Forest Road 1177, 14 mi N and 22 mi E Sweet Home, Willamette National Forest , Linn County, Oregon. North American distribution: USA: OR.

P. borealis Peck and Cook, 2011: 18 . Holotype male in CMNC. Type locality: Chena Ridge , 5 mi W Fairbanks, Alaska. North American distribution: USA: AK; occurring north of the limits of Pleistocene glaciations as a probable relict of a wider Tertiary distribution.

P. capizzii ( Hatch, 1957: 21) (Catopocerus) ; Peck and Cook 2011: 41. Holotype male in USNM. Type locality: Humbug Mt. , Curry County, Oregon. North American distribution: USA: ID, OR, WA. Biology : Fogel and Peck (1975).

P. chandleri Peck and Cook, 2011: 22 . Holotype male in FMNH. Type locality: 6 mi W Log Springs, Mendocino National Forest , Tehama County, California. North American distribution: USA: CA.

P. colorado Peck and Cook, 2011: 20 . Holotype male in CMNC. Type locality: 10 mi N Wolcott, Eagle County, Colorado. North American distribution: USA: CO.

P. constrictus Peck and Cook, 2011: 23 . Holotype male in FMNH. Type locality: 2.0 km off on Forest Road 2N46, Angeles Crest Highway , Los Angeles County, California. North American distribution: USA: CA.

P. contortus Peck and Cook, 2011: 23 . Holotype male in CDAE. Type locality: 5 mi E Placerville on highway 50, El Dorado County, California. North American distribution: USA: CA.

P. cryptophagoides ( Mannerheim, 1852: 333) (Catops) ; Horn 1880: 249 ( Pinodytes ); Hatch 1928: 72, 1957: 20 ( Catopocerus ); Peck and Cook 2011: 38 (distribution restricted). Lectotype in MCZC (designated by Peck and Cook 2011: 38); paralectotype in MZHF (in Mannerheim coll.) ( Silfverberg 1987: 49). Type locality: “Sitkha” {Sitka], Alaska. North American distribution: Canada: BC. USA: AK, WA (misidentified from Washington, DC by Horn (1880: 249) and from California by ( Leng 1920: 87), teste Peck and Cook (2011)). Biology: Fogel and Peck (1975); Peck and Cook (2011).

P. delnorte Peck and Cook, 2011: 44 . Holotype male in CMNC. Type locality: 10 km E Crescent City, Jed Smith Redwood State Park , Del Norte County, California. North American distribution: USA: CA.

P. eldorado Peck and Cook, 2011: 24 . Holotype male in CDAE. Type locality: 1.6 mi W Quintette, El Dorado County, California. North American distribution: USA: CA.

P. fresno Peck and Cook, 2011: 25 . Holotype male in FMNH. Type locality: Tamarack Ridge, 3.4 mi SE Highway 168, Sierra National Forest , Fresno County, California. North American distribution: USA: CA.

P. garibaldi Peck and Cook, 2011: 45 . Holotype male in OSUO. Type locality: 2 mi N Garibaldi, Tillamook County, Oregon. North American distribution: USA: OR (not WA).

P. gibbosus Peck and Cook, 2011: 25 . Holotype male in CMNC. Type locality: 20 km SW Lake Elsinore, Route 74, Orange County, California. North American distribution: USA: CA.

P. haidagwaii Peck and Cook, 2011: 45 . Holotype male in CNCI. Type locality: Kiusta , Graham Island , Queen Charlotte Island, British Columbia. North American distribution: Canada: BC.

P. humboldtensis Peck and Cook, 2011: 27 . Holotype male in CMNC. Type locality: Orick , Lady Byrd Johnson Grove , Redwood National Park , Humboldt County, California. North American distribution: USA: CA.

P. idaho Peck and Cook, 2011: 28 . Holotype male in WFBM. Type locality: 10 mi E Slate Creek, Slate Creek Campground , Idaho County, Idaho. North American distribution: USA: ID.

P. imbricatus ( Hatch, 1957: 20) (Catopocerus) ; Peck and Cook 2011: 40. Holotype female in USNM. Type locality: 6 mi N Buxton, Washington County , Oregon. North American distribution: USA: OR.

P. isabella Peck and Cook, 2011: 46 . Holotype male in WFBM. Type locality: Isabella Creek , Clearwater County, Idaho. North American distribution: USA: ID.

P. klamathensis Peck and Cook, 2011: 28 . Holotype male in CMNC. Type locality: 18 km NE Crescent City, Myrtle Creek Botanical Reserve , Del Norte County, California. North American distribution: USA: CA, OR.

P. losangeles Peck and Cook, 2011: 29 . Holotype male in USNM. Type locality: No precise location, Los Angeles County, California. North American distribution: USA: CA.

P. marinensis Peck and Cook, 2011: 30 . Holotype male in FMNH. Type locality: Tocaloma , Marin County, California. North American distribution: USA: CA.

P. minutus Peck and Cook, 2011: 31 . Holotype male in FMNH. Type locality: 4.9 mi W Pine Grove, Amador County, California. North American distribution: USA: CA.

P. monterey Peck and Cook, 2011: 31 . Holotype male in CDAE. Type locality: 2.2 mi W Bottchers Gap, Monterey County, California. North American distribution: USA: CA.

P. newelli ( Hatch, 1957: 20) (Catopocerus) ; Peck and Cook 2011: 41. Holotype male in USNM. Type locality: Coos Head , Coos County, Oregon. North American distribution: USA: OR, WA.

P. newtoni Peck and Cook, 2011: 17 . Holotype male in UADE. Type locality: Queen Wilhelmina State Park , Polk County, Arkansas. North American distribution: USA: AR, MO, TX.

P. orca Peck and Cook, 2011: 47 . Holotype male in CMNC. Type locality: Oregon Cave , Oregon Caves National Monument , Josephine County, Oregon. North American distribution: USA: OR.

P. ovatus Hatch, 1957: 21 (Catopocerus) ; Peck and Cook 2011: 21. Holotype male in USNM. Type locality: 5 mi N Gold Beach, Oregon. Peck and Cook 2011: 21. North American distribution: USA: OR. Biology : Fogel and Peck (1975).

P. parvus Peck and Cook, 2011: 32 . Holotype male in CDAE. Type locality: 5 mi S Gasquet, Del Norte County, California. North American distribution: USA: CA.

P. punctatus Peck and Cook, 2011: 19 . Holotype male in FMNH. Type locality: Mt. Spokane , Spokane County, Washington. North American distribution: USA: ID, WA.

P. pusio Horn, 1892: 45 ; Hatch 1928: 72 ( Catopocerus ); Peck and Cook 2011: 21. Lectotype male in MCZC (designated by Peck and Cook 2011: 21). Type locality: Alameda County, California.North American distribution: USA: CA.

P. rothi Hatch, 1957: 21 (Catopocerus) ; Peck and Cook 2011: 42. Holotype male in USNM. Type locality: 6 mi N Buxton, Washington County , Oregon. North American distribution: USA: CA, OR. Biology : Fogel and Peck (1975).

P. sanjacinto Peck and Cook, 2011 ; 33. Holotype male in CMNC. Type locality: Lake Fulmor , 15 km NW Idyllwild, San Jacinto Mountains , Riverside County, California. North American distribution: USA: CA.

P. sequoia Peck and Cook, 2011: 34 . Holotype male in CMNC. Type locality: Carmoe Crevice Cave , Tulare County, California. North American distribution: USA: CA.

P. setosus Peck and Cook, 2011: 48 . Holotype male in FMNH. Type locality: 6.7 mi NE Brookings, Curry County, Oregon. North American distribution: USA: CA, OR.

P. shasta Peck and Cook, 2011: 35 . Holotype male in FMNH. Type locality Buckhorn Pass , Highway 299, Shasta-Trinity County line, California. North American distribution: USA: CA.

P. shoshone Peck and Cook, 2011: 48 . Holotype male in WFBM. Type locality: Hobo Cedar Grove , Shoshone County, Idaho. North American distribution: USA: ID.

P. sinuatus Peck and Cook, 2011: 49 . Holotype male in CMNC. Type locality: 7 mi S, 13 mi E, Ashland , Soda Mountain Road , Jackson County, Oregon. North American distribution: USA: OR.

P. spinus Peck and Cook, 2011: 36 . Holotype male in CDAE. Type locality: 5 mi NE Butte Meadows, Butte County, California. North American distribution: USA: CA.

P. subterraneus ( Hatch, 1935: 115) (Typhloleiodes) ; Hatch 1957: 19 ( Catopocerus ); Peck and Cook 2011: 39 (distribution restricted, not BC). Holotype male in USNM. Type locality: Seaside, Clatsop County, Oregon. North American distribution: USA: OR. Biology: Fogel and Peck (1975).

P. tehama Peck and Cook, 2011: 36 . Holotype male in DENH. Type locality: 5 mi SE Manton, Bluff Springs , Tehama County; California. North American distribution: USA: CA.

P. tibialis ( Hatch, 1957: 20) (Catopocerus) ; Peck and Cook 2011: 43. Holotype male in USNM. Type locality: Gold Beach , Curry County, Oregon. North American distribution: USA: OR.

P. tuolumne Peck and Cook, 2011: 37 . Holotype male in CMNC. Type locality: Harden Flat , Tuolumne County, California. North American distribution: USA: CA.