Cherevychnavis, Bochenski & Wertz & Tomek & Gorobets, 2019

Bochenski, Zbigniew M., Wertz, Krzysztof, Tomek, Teresa & Gorobets, Leonid, 2019, A new species of the late Miocene charadriiform bird (Aves: Charadriiformes), with a summary of all Paleogene and Miocene Charadrii remains, Zootaxa 4624 (1), pp. 41-58 : 43-53

publication ID	https://doi.org/10.11646/zootaxa.4624.1.3
publication LSID	lsid:zoobank.org:pub:84AA439F-5A44-495B-A981-81ED53173690
persistent identifier	https://treatment.plazi.org/id/842887D0-EF3E-813F-FF3E-FDFE3B7A17C3
treatment provided by	Plazi
scientific name	Cherevychnavis
status	gen. nov.

Treatment

Genus: CHEREVYCHNAVIS gen. nov.

Type species. Cherevychnavis umanskae sp. nov.

Etymology. The genus name is the name of the type locality ( Cherevychne) merged with the Latin word for bird ( avis). Gender is feminine.

Diagnosis. A member of Charadrii , approximately the size of extant Haematopus ostralegus that differs from all genera by the following combination of characters of the coracoid: robust processus acrocoracoideus with cranio-sternal length about the same as medial projection; the foramen nervi supracoracoidei is very close to the sternal margin of the cotyla scapularis (only in Haematopus is this foramen similarly close, yet is still more widely separated); the medial margin of the facies articularis humeralis is straight almost along its entire length; and sternally, the angulus lateralis is broad with sternal and lateral margins meeting almost at right angles; and on which the dorsal edge of the facies articularis sternalis connects to the angulus lateralis.

Differential diagnosis. The coracoid and humerus differ as follows:

From Haematopus , Recurvirostra and Himantopus in:

Coracoid ( Figs. 2 View FIGURE 2 –4 View FIGURE 2 View FIGURE 3 View FIGURE 4 ): ● extremitas omalis, dorsal view: The medial margin of the facies articularis humeralis is straight almost along its entire length (#5) (in H. ostralegus , Recurvirostra and Himantopus it arches or curves medially towards the facies articularis clavicularis); ● extremitas omalis, dorsal view: The foramen nervi su- pracoracoidei is very close to the sternal margin of the cotyla scapularis (#9) (in H. ostralegus , Recurvirostra and Himantopus the foramen is somewhat farther sternad from the cotyla); ● extremitas sternalis, dorsal view: The dorsal edge of the facies articularis sternalis connects to the angulus lateralis (#14) (in H. ostralegus , most Recurvirostra and Himantopus the dorsal edge of the facies articularis sternalis disappears well before reaching the angulus lateralis).

Humerus ( Figs. 9–10 View FIGURE 9 View FIGURE 10 ): ● caudalodistal view: The infraspinatus attachment on crista bicipitalis is triangular, very broad dorsoventrally (#8) (in H. ostralegus , it is elongated proximodistally and much narrower; in Recurvirostra it is roundish and in Himantopus it is irregular); ● dorsal view, cranial margin of caput humeri: The caput humeri is more rounded and does not reach far proximally (#10) (in H. ostralegus , Recurvirostra and Himantopus it is more elongated and reaches farther proximally); ● ventral view: There is a deep rounded depression in the crista bicipitalis immediately distal to the tuberculum ventrale (#9) (in H. ostralegus , Recurvirostra and Himantopus the depression is shallower and its edges are less conspicuous); ● cranial view: Proximal part of the impressio musculus coracobrachialis is deeper and its edges are better distinguished (#12) than in extant H. ostralegus , Recurvirostra and Himantopus .

From Pluvialis in:

Coracoid ( Figs. 2 View FIGURE 2 –4 View FIGURE 2 View FIGURE 3 View FIGURE 4 and 6 View FIGURE 6 ): ● slender with relatively thinner shaft (#10) (in Pluvialis the bone is stout with thick shaft); ● extremitas omalis, medial view: sternal margin of facies articularis clavicularis is slanting (#21) (in Pluvialis , it is almost perpendicular to the long axis of the bone); ● extremitas omalis: medial area in sulcus supracoracoideus adjacent to the facies articularis humeralis deeply concave (#6) (in Pluvialis , the area is flat or shallowly concave); ● extremitas omalis, dorsal view: the foramen nervi supracoracoidei is very close to the sternal margin of the cotyla scapularis (#9) (in Pluvialis , the foramen is distinctly farther sternad from the cotyla); ● extremitas sternalis, dorsal and/or ventral view: facies articularis sternalis is relatively long (wide) and almost straight lateromedially because angulus medialis and angulus lateralis do not protrude far sternally (#15) (in Pluvialis, facies articularis sternalis is relatively short lateromedially and significantly bent because angulus medialis and angulus lateralis protrude far sternally); ● extremitas sternalis, dorsal and/or ventral view: the angulus lateralis is broad with sternal and lateral margins meeting almost at right angles (#13) (in Pluvialis, it tapers and its edges meet at an acute angle). Humerus ( Fig. 8 View FIGURE 8 ): ● caudal view: proximal part of the capital shaft ridge curves ventrad before reaching the caput humeri (#4) (in Pluvialis, the ridge is straight); ● tuberculum dorsale relatively short proximodistally (#1) (in Pluvialis, it is relatively longer).

From Charadrius in:

Coracoid ( Figs. 3 View FIGURE 3 and 5–6 View FIGURE 5 View FIGURE 6 ): ● extremitas omalis, ventral view: the processus acrocoracoideus is robust, with cranio-sternal length about the same as medial projection (#16) (in Charadrius , the width of the processus acrocoracoideus is clearly greater than its cranio-sternal length); ● extremitas omalis, ventral view: the ligamental scar on the acrocoracoid (for attachment of ligamentum acrocoracoclaviculare superficiale, Worthy et al. 2009, De Pietri et al. 2018) is shallow and elongated (#17) (in Charadrius, it forms a deep, rounded, conspicuous depression); ● extremitas omalis: the recess below the facies articularis clavicularis, especially in its ventral part, is shallow (#22) (in Charadrius, the recess is very deep, also in its ventral part); ● extremitas omalis, dorsal view: the foramen nervi supracoracoidei is very close to the sternal margin of the cotyla scapularis (#9) (in Charadrius, the foramen is distinctly farther sternad from the cotyla).

Humerus ( Figs. 9–10 View FIGURE 9 View FIGURE 10 ): ● caudalodistal view: the infraspinatus attachment on crista bicipitalis is triangular, very broad dorsoventrally (#8) (in Charadrius , it is elongated proximodistally and much narrower); ● cranial view: the boundary between intumescentia and shaft is difficult to define; the transition is gentle, gradual (#14) (in Charadrius, the transition is clearly marked); ● ventral view: there is a deep rounded depression in the crista bicipitalis immediately distal to the tuberculum ventrale (#9) (in Charadrius, the depression is shallower and its edges are less conspicuous).

From Vanellus in:

Coracoid ( Figs. 3–5 View FIGURE 3 View FIGURE 4 View FIGURE 5 ): ● extremitas omalis, ventral view: the processus acrocoracoideus is robust, with cranio-sternal length about the same as medial projection (#16) (in Vanellus , the width of the processus acrocoracoideus is clearly greater than its cranio-sternal length); ● extremitas omalis, dorsal view: cotyla scapularis very large, its lateral margin protruding laterally beyond the lateral edge of the bone (#8) (in Vanellus , cotyla scapularis relatively small, its lateral margin does not extend laterally beyond the lateral edge of the bone); ● extremitas omalis, dorsal view: the foramen nervi supracoracoidei is very close to the sternal margin of the cotyla scapularis (#9) (in Vanellus, the foramen is distinctly farther sternad from the cotyla); ● extremitas sternalis, dorsal view: the bone surface above the articulation is flat or only slightly concave (#12) (in Vanellus, there is a deep depression there); ● extremitas sternalis: there is a bony blade just above the projection on the margo medialis (#11) (in Vanellus, the blade is missing); ● extremitas sternalis, dorsal and/or ventral view: facies articularis sternalis is long (wide) and almost straight lateromedially because angulus medialis and angulus lateralis do not protrude far sternally (#15) (in Vanellus, facies articularis sternalis is relatively short lateromedially and significantly bent because angulus medialis and angulus lateralis protrude far sternally).

Humerus ( Figs. 8 View FIGURE 8 –10 View FIGURE 8 View FIGURE 9 View FIGURE 10 ): ● caudal view: proximal part of the capital shaft ridge curves ventrad before reaching the caput humeri (#4) (in Vanellus , the ridge is straight); ● caudalodistal view: caput humeri in its ventral part is moderately deeply undercut (#5) (in Vanellus , caput humeri is very deeply undercut); ● caudalodistal view: the infraspinatus attachment on crista bicipitalis is triangular, very broad dorsoventrally (#8) (in Vanellus, it is elongated proximodistally and much narrower); ● cranial view: the boundary between intumescentia and shaft is difficult to define; the transition is gentle, gradual (#14) (in Vanellus, the transition is clearly marked); ● the proximal part of the crista pectoralis slopes dorsally (#3) (in Vanellus, it is almost parallel to the long axis of the bone); ● ventral view: there is a deep rounded depression in the crista bicipitalis immediately distal to the tuberculum ventrale (#9) (in Vanellus, the depression is shallower and its edges are less conspicuous).

From Pluvianus in:

Coracoid ( Figs. 2 View FIGURE 2 –5 View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 ): ● extremitas omalis: the section between processus procoracoideus and processus acrocora- coideus is elongated (#1) (in Pluvianus , it is short in relation to the total length of the coracoid); ● extremitas omalis, ventral view: the processus acrocoracoideus is robust, with cranio-sternal length about the same as medial projection (#16) (in Pluvianus , the width of the processus acrocoracoideus is clearly greater than its cranio-sternal length); ● extremitas omalis, dorsal view: the foramen nervi supracoracoidei is very close to the sternal margin of the cotyla scapularis (#9) (in Pluvianus , the foramen is distinctly farther sternad from the cotyla); ● extremitas sternalis, dorsal view: the dorsal edge of the facies articularis sternalis connects to the angulus lateralis (#14) (in Pluvianus , the dorsal edge of the facies articularis sternalis disappears well before reaching the angulus lateralis); ● extremitas sternalis, dorsal and/or ventral view: the angulus lateralis is broad with sternal and lateral margins meeting almost at right angles (#13) (in Pluvianus , it tapers and its edges meet at an acute angle); ● extremitas sternalis: the projection on the margo medialis is small, dull and ends little above the angulus medialis (#18) (in Pluvianus , it is bigger, sharp and reaches high above angulus medialis).

Humerus ( Figs. 9–10 View FIGURE 9 View FIGURE 10 ): ● caudalodistal view: the infraspinatus attachment on crista bicipitalis is triangular, very broad dorsoventrally (#8) (in Pluvianus , it is elongated proximodistally and much narrower); ● cranial view: the proximal part is approximately as long proximodistally as it is broad dorsoventrally (#11) (in Pluvianus , the proximal part is shorter proximodistally); ● cranial view: the boundary between intumescentia and shaft is difficult to define; the transition is gentle, gradual (#14) (in Pluvianus , the transition is clearly marked).

From Burhinus in:

Coracoid ( Figs. 2 View FIGURE 2 –3 View FIGURE 2 View FIGURE 3 and 5–6 View FIGURE 5 View FIGURE 6 ): ● extremitas omalis: the lateral edge of processus acrocoracoideus is oblique to the long axis of the bone (#4) (in Burhinus , the lateral edge is more vertical); ● extremitas omalis, ventral view: the processus acrocoracoideus is robust, with cranio-sternal length about the same as medial projection (#16) (in Burhinus , the width of the processus acrocoracoideus is clearly greater than its craniosternal length);● extremitas omalis, medial view: sternal margin of facies articularis clavicularis is broad ventrodorsally (#20) (in Burhinus , it is relatively narrow); ● extremitas omalis, dorsal view: the foramen nervi supracoracoidei is very close to the sternal margin of the cotyla scapularis (#9) (in Burhinus , the foramen is distinctly farther sternad from the cotyla); ● extremitas sternalis, ventral view: the bone surface, medial to the linea intermuscularis ventralis, is flat or slightly convex (#19) (in Burhinus , there is a distinct depression there); ● extremitas sternalis: the projection on the margo medialis is small, dull and ends little above the angulus medialis (#18) (in Burhinus , it is bigger, sharp and reaches high above angulus medialis).

Humerus ( Figs. 8 View FIGURE 8 and 10 View FIGURE 10 ): ● caudal view: a well-developed dorsal (second) fossa pneumotricipitalis is present (#6) (in Burhinus , the dorsal fossa is absent); ● caudal view: the proximal edge of tuberculum dorsale slopes gently distally (#2) (in Burhinus , tuberculum dorsale is more pronounced and its proximal edge is often upturned proximally); ● caudal view: crista bicipitalis arches evenly (#7) (in Burhinus , the infraspinatus attachment on crista bicipitalis protrudes clearly more ventrally); ● cranial view: impressio musculus coracobrachialis relatively narrow (#13) (in Burhinus , it is very wide, reaching at least half the width of the proximal humerus).

From Chionis in:

Coracoid ( Figs. 2 View FIGURE 2 –5 View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 ): ● extremitas omalis: the section between processus procoracoideus and processus acrocoracoideus is elongated (#1) (in Chionis , it is short in relation to the total length of the coracoid); ● extremitas omalis, ventral view: the ligamental scar on the acrocoracoid (for attachment of ligamentum acrocoracoclaviculare superficiale, Worthy et al. 2009, De Pietri et al. 2018) is shallow and elongated (#17) (in Chionis, it forms a deep, rounded, conspicuous depression); ● extremitas omalis, ventral view: processus acrocoracoideus protrudes far medially (#7) (in Chionis, the processus protrudes very little medially); ● extremitas omalis, omal view: processus acrocoracoideus is dorsoventrally narrow (#2) (in Chionis, it is very broad); ● extremitas omalis, omal view: ventral edge is slightly bent (#3) (in Chionis, ventral edge is strongly arched because facies articularis humeralis and facets articularis clavicularis protrude far ventrally); ● extremitas omalis, dorsal view: the foramen nervi supracoracoidei is very close to the sternal margin of the cotyla scapularis (#9) (in Chionis, the foramen is distinctly farther sternad from the cotyla); ● extremitas sternalis, dorsal and/or ventral view: facies articularis sternalis is long (wide) and almost straight lateromedially because angulus medialis and angulus lateralis do not protrude far sternally (#15) (in Chionis, facies articularis sternalis is relatively short lateromedially and significantly bent because angulus medialis and angulus lateralis protrude far sternally).

Humerus ( Figs. 9–10 View FIGURE 9 View FIGURE 10 ): ● caudalodistal view: The infraspinatus attachment on crista bicipitalis is triangular, very broad dorsoventrally (#8) (in Chionis , it is elongated proximodistally and much narrower); ● dorsal view, cranial margin of caput humeri: The caput humeri is more rounded and does not reach far proximally (#10) (in Chionis , it is more elongated and reaches farther proximally); ● ventral view: There is a deep rounded depression in the crista bicipitalis immediately distal to the tuberculum ventrale (#9) (in Chionis , the depression is shallower and its edges are less conspicuous); ● cranial view: Proximal part of the impressio musculus coracobrachialis is deeper and its edges are better distinguished (#12) than in most extant Chionis .