Syzygium bullatum (Brongn. & Gris) N. Snow & Byng, 2017

Snow, Neil, Byng, James W., Munzinger, Jérôme, Callmander, Martin W. & Dawson, John W., 2017, New Caledonian Piliocalyx transferred to Syzygium (Myrtaceae) with an updated conspectus of the species, Candollea 72 (2), pp. 239-248 : 241-243

Syzygium bullatum (Brongn. & Gris) N. Snow & Byng , comb. nov.

Ξ Piliocalyx bullatus Brongn. & Gris in Bull. Soc. Bot. France 13: 470. 1866.

Lectotypus (designated here): NEW CALEDONIA. Prov. Nord: Wagap, Vieillard 2177 ( P [ P00543883 ]!; isolecto-: P [ P00543884 , P00543885 ]!) . Syntypi: NEW CALEDONIA : Prov. Nord : Wagap, Vieillard 2177 ( A [A 00071209] image seen ; G [ G00341008 ]!; GH [ GH00071208 ] image seen, K [ K000800626 ]!; Z [ Z-000056172 ]!) .

Trees 7- 30 m. Bark grayish, mostly smooth. Branchlets terete, becoming grayish. Leaves sessile. Leaf blades c. 7-22 × 4.5-6.0 cm, oblong to narrowly oblong or narrowly ovate, base cordate to subcordate, margin revolute, apex obtuse, venation moderately to conspicuously bullate. Inflorescences terminal, paniculate, multi-branched, 5-19 × 4-8 cm (sometimes c. 2 × length of subtending leaves); axes laterally compressed beneath branching points, light yellowish green. Hypanthium 1.8-2 × c. 2.5 mm, whitish; filaments whitish to magenta. Fruits 1.2-2.2 × 1.3-2.3 cm; globular to subglobular, base rounded to somewhat tapered, maturing whitish, cream-colored or yellowish.

[B: Munzinger & Dagostini 2167; C: Munzinger 5600; D: Munzinger 2221; E: Munzinger 4915; F: Munzinger 2416]

[Photos: A-B, D-F: J. Munzinger; C: P. Lowry]

Notes. ‒ The protologue indicated only Vieillard 2176 from Wagap, which is an error because the marked (“type”) specimen clearly indicates 2177, and because 2176 is listed from the same locality as the type of Piliocalyx wagapensis Brongn. & Gris in the same publication on the next page. The specimen marked with a “type” tag at P for Syzygium bullatum (Vieillard 2177) , selected here as the lectotype, is indisputably representative of that taxon. Specimens of the same number are at G, K, and Z but these bear different labels. We thus cannot be certain that these remaining syntypes are of the same gathering, which was made when the concept of types did not exist as we know it today. We thus follow a conservative treatment in designating a lectotype with remaining syntypes rather than citation of a lectotype with isolectotypes ( MCNEILL, 2014).

The leaf blades of McPherson 5090 south of col d’Amieu are markedly narrower than other specimens, but other diagnostic aspects of the leaf (e.g., bullate texture, widely diverging and spacing of secondary veins) and the whitish fruit accord with our concept of this species.

A specimen at P previously labeled as this taxon (MacKee 13401) is S. dawsonianum N. Snow, S.L. Young & Callm. ( SNOW et al., 2016) . Comments concerning fresh material (above) are based in part on a color photograph that was sent to Odile Poncy (MNHN) formerly in the possession of Craven, which shows a mature flowering specimen of S. bullatum , although it is neither labeled as such nor refers to a voucher.

Additional specimens examined. ‒ NEW CALEDONIA. Prov. Nord: Poami (the Temala), 1.IX.2010, Hequet et al. 3880 ( NOU [ NOU080968 ]) ; Haute Amoa, 26.XII.1968, MacKee 20048 ( NOU [ NOU013794 ], P [ P04776435 , P05265291 , P05265299 ]) ; Poindimié , 14.V.1973, MacKee 26701 ( NOU [ NOU013791 ], P [ P00500688 ]) ; Tiwaka , Moindi , 31.VI.1978, MacKee 35228 ( NOU [ NOU013790 ], P [ P05265221 ]) ; Aoupinié , 15.X.2008, Munzinger et al. 5191 ( NOU [ NOU050169 ]) ; Mandjélia , 10.I.2009, Munzinger et al. 5842 ( NOU [ NOU053515 ], P [ P06669059 ]) ; La Guen , 20.XI.2010, Munzinger et al.6384 ( NOU [ NOU063546 ], P [ P00805948 ]) . Prov. Sud: Crête au N du col d’Amieu , 24.IX.1965, MacKee 13471 ( NOU [ NOU013793 ], P [ P04633434 ]) ; col d’Amieu , X.1979, MacKee 37447 ( P [ P06603439 , P04633434 ]) ; near Mt. Rembai , south of Col d’Amieu , 7.XI.1982, McPherson 5090 ( MO-3233405 , NOU [ NOU013795 ], P [ P04884357 ]) .