Bembidion quadrimaculatum dubitans (LeConte, 1852)

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 385-386

Bembidion quadrimaculatum dubitans (LeConte, 1852)

Ochthedromus dubitans LeConte, 1852a: 189. Type locality: "San Francisco [San Francisco County, California]" (original citation). Syntype(s) in MCZ [# 5548].

Ochthedromus cruralis LeConte, 1852a: 189. Type locality: "San Jose [Santa Clara County, California]" (original citation). Syntype(s) in MCZ [# 5549]. Synonymy established by Hayward (1897: 136), confirmed by Lindroth (1963b: 384).

Bembidion gregale Casey, 1918: 148. Type locality: "Agassiz, British Columbia" (original citation). Lectotype (♀), designated by Lindroth (1975: 121), in USNM [# 37060]. Synonymy established by Hatch (1953: 98), confirmed by Lindroth (1963b: 384).

Bembidion pugetanum Casey, 1918: 148 [primary homonym of Bembidion pugetanum Fall, 1916]. Type locality: "Spokane [Spokane County], Washington" (original citation). Lectotype (♂), designated by Lindroth (1975: 121), in USNM [# 37061]. Synonymy established by Hatch (1953: 98), confirmed by Lindroth (1963b: 384).

Bembidion sapphicum Casey, 1918: 149. Type locality: "Reno [Washoe County], Nevada" (original citation). Lectotype (♂), designated by Lindroth (1975: 121), in USNM [# 37063]. Synonymy established by Lindroth (1963b: 384).

Bembidion tenax Casey, 1918: 152. Type locality: "Fort Wingate [McKinley County], New Mexico" (original citation). Lectotype (♂), designated by Erwin (1984a: 180), in USNM [# 37065]. Synonymy established by Erwin (1984a: 180).

Bembidion caseyi Leng, 1919b: 202. Replacement name for Bembidion pugetanum Casey, 1918.

Distribution.

This subspecies ranges from the southwestern edge of Lac Winnipeg in southern Manitoba to Vancouver Island (Hayward 1897: 119), north to the Brook Range in Alaska (Lindroth 1963b: 385), south to the Baja California Peninsula (Horn 1894: 308) and southern New Mexico (Fall and Cockerell 1907: 157). The records from Kansas (Snow 1903: 193; Knaus 1905a: 218) need confirmation.

Records.

CAN: AB, BC (VCI), MB, NT, SK, YT USA: AK, AZ, CA, CO, ID, MT, NM, NV, OR, WA, WY [KS] - Mexico

Note.

Three other subspecies, including the nominotypical one, are known from the Palaearctic Region (Toledano 1999: 205-210).