Nanometa tetracaena, Álvarez-Padilla & Kallal & Hormiga, 2020

Álvarez-Padilla, Fernando, Kallal, Robert J. & Hormiga, Gustavo, 2020, Taxonomy And Phylogenetics Of Nanometinae And Other Australasian Orb-Weaving Spiders (Araneae: Tetragnathidae), Bulletin of the American Museum of Natural History 2020 (438), pp. 1-107: 1-107

publication ID

http://doi.org/ 10.1206/0003-0090.438.1.1

DOI

http://doi.org/10.5281/zenodo.4631653

persistent identifier

http://treatment.plazi.org/id/881F3552-7616-A333-FF54-6AFFFEF1FD75

treatment provided by

Felipe

scientific name

Nanometa tetracaena
status

sp. nov.

Nanometa tetracaena   , sp. nov.

Figures 18–19 View FIGURE 18 View FIGURE 19 , 37 View FIGURE 37

TYPE MATERIAL: Male holotype and female allotype from Australia, Victoria, Strzelecki Ranges, Gunyah-Toora Rd. , 2 km SSW of Gunyah, -38.541, 146.317, G. Milledge, 5 March 1995 ( NMV K-6309) GoogleMaps   ; both types deposited at NMVM.

DIAGNOSIS: Males of N. tetracaena   are most likely to be confused with N. dutrorum   . The CEMP of N. tetracaena   (figs. 18D, 19C) does not bear the conspicuous spine found in N. dutrorum   (figs. 31C, 33C), its conductor lacks the serrated denticles found in N. dutrorum   (fig. 33A–C), and the embolic process is more sclerotized than that of N. dutrorum   (fig. 31A, E). The CEBP of N. tetracaena   has four apophyses, whereas N. dutrorum   has three. The epigynum of N. tetracaena   has a small carina separating the copulatory openings (figs. 18E, 19D), which is readily distinguishable from the shallow, M-shaped ridge on the epigynum of N. dutrorum   .

DESCRIPTION: Female (NMV K-6295) total length 3.14. Cephalothorax length 1.09, width 0.85. Clypeus height 1.0 AME diameter. Cephalothorax background pale yellow, with light gray concentrated over ocular area, carapace lateral edges, and near margin of pars cephalica and pars thoracica (fig. 18A). Endites and sternum grayish yellow (fig. 18C). Cheliceral promargin and retromargin with three and two teeth, respectively, cheliceral denticles present. Abdomen dorsum background pale brown, silvery guanine patches covering all abdomen dorsal surface, except a medial line that extends the entire abdomen length (fig. 18A), lateral surface covered with guanine patches over dorsal half, ventral half with brown lines (fig. 18A, C), ventral surface brown intercalated with silvery guanine patches and with a central rectangle flanked by two lines (fig. 18C). Booklung stridulatory organ, tracheae and epigynum not observed with SEM. Femur I length 1.7.

Male (NMV K-6305) same as female except as noted. Total length 2.75. Cephalothorax length 1.15 width 0.94. Clypeus 1.5 AME diameter. Cephalothorax and abdomen coloration slightly darker than in female, silvery guanine patches less conspicuous and reduced into longitudinal stripes. Cheliceral promargin and retromargin with three and two teeth respectively. Cheliceral denticles present, stridulatory organ and coxa IV retrolateral surface not observed with SEM. Femur I length 2.6. Palp with conspicuous CEMP and CEBP with four pointed apophyses.

VARIATION: Females (N = 3) total length 2.82– 3.27, cephalothorax length 1.13–1.27, width 0.84–0.94. Males (N = 3) total length 2.41–2.82, cephalothorax length 0.99–1.30, width 0.82–1.08.

ETYMOLOGY: The specific epithet is from the Greek tetra (four) and acaena (spikes or thorns), referring to the four points on the CEBP.

DISTRIBUTION: Nanometa tetracaena   can be found in Tasmania, Victoria and southern New South Wales (fig. 37D).

MATERIAL EXAMINED: N = 88. AUSTRALIA, Victoria: Central Highlands, Acheron Gap , 6 km NE of Mt. Donna Buang, 37° 41′ 20.4″ S, 145° 44′ 20.4″ E, G. Milledge, 28 December 1995, Nothofagus cunninghami   forest, intercept trap, 3 males, 7 females, NMV K-6297 GoogleMaps   ; Central Highlands, Acheron Gap , 6 km NE of Mount Donna Buang, 37° 41′ 20.4″ S, 145° 44′ 20.4″ E, G. Milledge, 20 February 1996, direct search, 2 males, NMV K-6317 GoogleMaps   ; Central Highlands, Acheron Gap, 6 km NE of Mt. Donna Buang, 37° 41′ 20.4″ S, 145° 44′ 20.4″ E, G. Milledge, 28 December 1995 – 21 February 1996, direct search, GoogleMaps   1 male, 3 females, NMV K-6298. Central Highlands , The Big Culvert , 2.5 km ENE of Mt. Observation , 37° 33′ 36″ S, 145° 52′ 15.6″ E, G. Milledge, 19 February 1996, Nothofagus cunninghami   forest, direct search, 2 males, 3 females, NMV K-6302 GoogleMaps   ; Mount Donna Buang, A. Neboiss, 5 February 1995, 1 male, 1 female, NMV   ; Great Otway National Park, Triplet Falls track, 38° 40′ 12″ S, 143° 29′ 52.80″ E, G. Hormiga & N. Scharff, 312 m, 16 April 2014, sifted litter, GWU ( GH1697) GoogleMaps   ; Otway Ranges, Aire Crossing Track, 0.5 km N of Aire R. Crossing , 38° 42′ 0″ S, 143° 28′ 58.8″ E, P. Lillywhite, 31 January 1995, Nothofagus cunninghami   forest, sweep net, 2 males, 3 females, NMV K-6293 GoogleMaps   ; Otway Ranges, Beauchamp Falls , 38° 38′ 60″ S, 143° 35′ 60″ E, G. Milledge, 15 November 1994 – 31 January 1995, intercept trap, 1 male, 2 females, NMV K-6830 GoogleMaps   ; Otway Ranges, Beauchamp Falls , 38° 38′ 60″ S, 143° 35′ 60″ E, G. Milledge, 31 January 1995 – 11 April 1995, 1 male, 2 females, NMV K-6294 GoogleMaps   ; Otway Ranges, Young’s Creek Rd, 0.4 km NW of Triplet Falls, 38° 40′ 1.2″ S, 143° 22′ 58.8 E, G. Milledge, 31 January 1995, Eucalyptus   sp. forest, intercept trap, 3 males, NMV K-6826 GoogleMaps   ; Otway Ranges, Young’s Creek Rd,. 0.4 km NW of Triplet Falls, 38° 40′ 1.2″ S, 143° 22′ 58.8 E, G. Milledge, 31 January 1995, Eucalyptus   sp. forest, direct search, 3 females, NMV K-6823 GoogleMaps   ; Otway Ranges, Young’s Creek Rd,. 0.4 km NW of Triplet Falls, 38° 40′ 1.2″ S, 143° 22′ 58.8 E, P. Lilywhite, 5 females, NMV K-6295 GoogleMaps   ; Phillips Track, Young’s Creek crossing, 0.5 km N of Triplet Falls, G. Milledge, 20 February 1992, Nothofagus cunninghami   forest, direct search, 2 females, 3 males, NMV   ; Strzelecki Ranges, Gunyah-Toora Rd. , 2 km SSW of Gunyah, 38° 32′ 31.2″ S, 146° 19′ 1.2″ E, G. Milledge, 5 March 1995, Nothofagus cunninghami   forest, intercept trap, 1 male, 1 female, NMV K-6386 GoogleMaps   ; Strzelecki Ranges, Gunyah-Toora Rd. , 2 km SSW of Gunyah, 38° 32′ 31.2″ S, 146° 19′ 1.2″ E, G. Milledge, 5 March 1995, Nothofagus cunninghami   forest, intercept trap, 1 male, 1 female, NMV K-6308 GoogleMaps   ; Strzelecki Ranges, Gunyah-Toora Rd. , 2 km SSW of Gunyah, 38° 32′ 31.2″ S, 146° 19′ 1.2″ E, G. Milledge, 5 March 1995, Nothofagus cunninghami   forest, intercept trap, 2 males, 12 females, NMV K-6309 GoogleMaps   ; Strzelecki Ranges, Gunyah-Toora Rd. , 2 km SSW of Gunyah, 38° 32′ 31.2″ S, 146° 19′ 1.2″ E, G. Milledge, 5 March 1995, Nothofagus cunninghami   forest, intercept trap, 3 males, NMV K-6305 GoogleMaps   ; Strzelecki Ranges, Tarra-Bulga National Park, 0.5 km NNE of Tarra Valley Picnic Area, 38° 26′ 38.4″ S. 146° 32′ 31.2″ E, G. Milledge, 10 January 1996 – 5 March 1996, Nothofagus cunninghami   forest, intercept trap, GoogleMaps   2 females, NMV K-6314. Tasmania: Mt. Michael , rainforest between Mt. Michael and Little Mt. Michael , 41° 10′ 58.8″ S, 148° 0′ 0.0″, R. Coy, P. Lillywhite, & A.L. Yen, 20 February 1990, Nothofagus cunninghami   forest, 2 males, 3 females, NMV GoogleMaps   ; Pirates Rd., 2.5 km SW of Eaglehawk Neck, 43° 3′ 10.8″ S, 147° 54′ 18″ E, R. Coy, P. Lillywhite, & A.L. Yen, 13 February 1990, Nothofagus cunninghami   forest, GoogleMaps   2 males, 3 females, NMV. New South Wales: Blue Mountains National Park, Mt. Wilson , 33° 29’ 55.75″ S, 150° 24’ 53.063″ E, 955 m, G. Hormiga & N. Scharff, 3 April 2014, 1 male, 1 female ( GH1662), GWU GoogleMaps   ; Jamberoo Mountain, 1 January 1993, J. Noble, 1F, KS 54234. Mt. St. Leonard , 37° 33′ 59.76″ S, 145° 31′ 59.88″ E, M.S. Harvey & M.E. Blosfelds, June 4 1991, 1 male, WAM T23237 View Materials GoogleMaps   ; Jamberoo Mountain, 1 January 1993, J. Noble, 1F, KS 54234. Mt. St. Leonard , 37° 33′ 59.76″ S, 145° 31′ 59.88″ E, M.S. Harvey & M.E. Blosfelds, June 4 1991, many females, WAM T23193 View Materials GoogleMaps   ; Coranderrk Reserve, Healesville, -37.7166, 145.5500, P.J. Gullan, GoogleMaps   1 male, WAM T24693 View Materials . Strathgordon , 42° 46′ 1.2″ S, 146° 3′ 0.0″ E, 6 January 1998, L.J. Boutin, 2MM, 2FF, QVM 13:43581 GoogleMaps   ; Mt. Wedge Track , 42° 52′ 58.8″ S, 146° 18′ 0.0″ E, 7 January 1998, L.J. Boutin, 1FF, 1M, QVM 13: 24035 GoogleMaps   ; Maggs Mountain , Eucalyptus   foliage, 9 December 1975, R.H. Green, 3FF, QVM 13: 44509   ; Maggs Mountain , 1979, R.H. Green, 1M, KS 52097   ; Strahan, rainforest nr. Hogarth Falls , 42° 8′ 60″ S, 145° 19′ 1.2″ E, 12 November 1997, L.J. Boutin, sweeping, 4MM, 1F, QVM 13:24059 GoogleMaps   ; Edgar Dam Campground, nr. Lake Peddar , 43° 1′ 58.8″ S, 146° 20′ 60″ E, 5 January 1998, L.J. Boutin, from sweeping, 2MM, 2FF, QVM 13:43593 GoogleMaps   ; Frenchmans Cap track, 42° 4′ 58.8″ S, 145° 55′ 58.80″ E, 24 December 1997, L.J. Boutin, mixed forest, sweeping, 2FF, 3MM, QVM 13:24066 GoogleMaps   .

NMV

Museum Victoria

WAM

Western Australian Museum