Enicospilus flavicaput (Morley, 1912)

Shimizu, So, 2020, The Nepalese species of the genus Enicospilus Stephens, 1835 (Hymenoptera, Ichneumonidae, Ophioninae): a preliminary revision and identification key to species, Deutsche Entomologische Zeitschrift 67 (1), pp. 69-126 : 69

Enicospilus flavicaput (Morley, 1912) Fig. 8

Enicospilus xanthocephalus Cameron 1907: 178; holotype ♀, Myanmar, NHMUK, examined; junior primary homonym of Enicospilus xanthocephalus Cameron, 1905.

Henicospilus flavicaput Morley 1912: 45; replacement name for Enicospilus xanthocephalus Cameron, 1907.

Henicospilus urospilus Enderlein 1921: 27; holotype ♀, Sumatra, IZPAN; synonymised by Townes et al. (1961: 72).

Material examined.

5♀♀ and 1 unsexed: Brunei (3♀♀), Indonesia (1♀), Myanmar (1♀), Sri Lanka (1 unsexed); no Nepalese specimens were examined .

Type series: holotype of Enicospilus xanthocephalus Cameron , 1907 (= Henicospilus flavicaput Morley, 1912), ♀, Haundraw Valley, Tenasserim, Myanmar, VIII.1894, C.T. Bingham leg. (NHMUK, Type 3b.1233) .

Non-type series: 1♀, U. Temburong (1,500 m), Bukit Retak , Brunei, IV.1981, I.D. Gauld leg. (Fig. 8 ) ; 1♀, Montane forest (1,618 m), Bukit Retak, Brunei, V.1979, I.D. Gauld leg. ; 1♀, Pagon Ridge , Pagon, Brunei, II.1982, G. Allen leg. ; 1♀, Perliawatte (1,200 -1,500 m), Mt Gede, West Java, Indonesia, I.1938 ; 1 unsexed, near Mahiyangana , Badulla Dist., Sri Lanka, 24.V.1974, Gans & Prasanna leg. (all NHMUK).

Distribution.

Australasian and Oriental regions ( Yu et al. 2016). Gauld and Mitchell (1981) recorded this species from Nepal.

Diagnosis.

Head (Fig. 8B-D ): GOI = 2.9-3.1; lower face 0.6-0.7 × as wide as high; clypeus weakly convex in profile, its lower margin subacute; mandible moderately twisted by 30-40°, moderately long, evenly tapered, its outer surface without a diagonal structure; upper mandibular tooth 1.3-1.6 × as long as lower one; posterior ocellus close to eye; antenna with 71-76 flagellomeres and 20th flagellomere 2.3-2.5 × as long as wide.

Mesosoma (Fig. 8E ): mesopleuron rather coarsely longitudinally striate; scutellum with lateral longitudinal carinae reaching anterior 0.8-1.0 and convergent posteriorly; metapleuron rather coarsely striate to strigose; propodeum evenly rounded to slightly declivous, its posterior area coarsely reticulate, outer margin of propodeal spiracle joining pleural carina by a strong ridge.

Wings (Fig. 8F ): fore wing with AI = 0.3-0.4, CI = 0.2-0.4, ICI = 0.6-0.7, SDI = 1.2-1.4; fore wing vein 1m-cu&M weakly sinuous, 2r&RS almost straight; fenestra and sclerites of discosubmarginal cell of fore wing as in Figure 8F ; fenestra of fore wing not long and its anterodistal corner distinctly separated from proximal end of vein RS; proximal sclerite triangular, confluent with distal one, strongly pigmented; central sclerite strongly pigmented and sclerotised, linear and parallel to vein 2r&RS, positioned in anterodistal part of fenestra; distal sclerite present proximally and vestigial to absent distally; proximal corner of marginal cell of fore wing uniformly setose; vein 1cu-a antefurcal to M&RS by 0.1-0.2 × 1cu-a length.

Colour (Fig. 8 ): body including interocellar area entirely testaceous; wings hyaline to weakly infuscate.

Differential diagnosis.

Enicospilus flavicaput is most similar to E. kanshirensis but can be distinguished from it by the slender central sclerite (Fig. 8F ) (central sclerite stouter in E. kanshirensis , as in Figure 14F ), and larger body size (i.e. fore wing length more than 17.0 mm in E. flavicaput but less than 15.0 mm in E. kanshirensis ).

Figures

Figure 8. Enicospilus flavicaput (Morley, 1912), ♀. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing.

Figure 14. Enicospilus kanshirensis (Uchida, 1928), ♂. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing.