Burmanteon olmii Engel, 2003

Xu, Zaifu, Olmi, Massimo & He, Junhua, 2013, Dryinidae of the Oriental region (Hymenoptera: Chrysidoidea), Zootaxa 3614 (1), pp. 1-460 : 200-201

1. Burmanteon olmii Engel, 2003

( Plate 52A – H )

Burmanteon olmii Engel 2003: 617 .

Description of female ( Plate 52A – H ). Fully winged; length 1.6 mm. Colour apparently black, except palpi, antenna and sting testaceous; clubs of femora with brown spot. Antenna 10-segmented, short and slender, covered with dense and short hairs, slightly thickened distally; antennal rhinaria apparently absent; antennal segments in following proportions: 5:2:7:5.5:5.5:5:4.5:4.5:4.5:6; antenna much shorter than body (1.2: 1.6 mm), approximately 2.5 times as long as head (length of head dorsally measured from occipital carina behind ocelli to distal apex of mandible): 49:20. Head slightly convex, with sculpture indistinct; mandible with four teeth progressing larger from anterior one to posterior. Face of holotype apparently crushed, appearing excavated (crushing regarding also anterior third of eyes); clypeus, face and frontal line not visible; occipital carina apparently complete, laterally not reaching eyes; occiput very excavated; eye normally bulging; POL = 2; OOL = 7; OL and TL not visible. Palpal formula 6/3. Maxillary and labial palpi covered with many long hairs. Pronotum long, crossed by anterior strong transverse impression, with posterior disc, without posterior collar. Pronotum covered with long hairs, with sculpture not distinct, slightly shorter than head (15:20), with posterior surface swollen and disc flat; posterior surface longer than anterior collar (10:5); pronotal tubercle apparently reaching tegula. Shape of pronotum similar to that of Lonchodryinus . Scutum difficult to discern, with long hairs, with sculpture not distinct, much shorter than pronotum (8:15). Notauli apparently complete, posteriorly separated; minimum distance between notauli difficult to discern, apparently approximately as long as antennal segment 2. Scutellum and metanotum difficult to discern. Scutellum + metanotum shorter than scutum (8:15). Propodeum approximately as long as scutum (16:15), with sculpture difficult to discern, but apparently reticulate rugose; posterior surface not visible; dorsal surface about as long as posterior surface. Mesopleura, metapleura and epicnemium difficult to discern. Shape of head, scutum, scutellum, metanotum and propodeum usual for Anteoninae . Fore wing ( Plate 52E, G ) hyaline, with usual venation of Anteoninae . Pterostigma long and narrow, much longer than broad (12:2). Shape of pterostigma similar to that of Dryininae . Marginal cell open. Stigmal vein regularly curved, with distal part much longer than proximal part (12:5); stigmal vein forming curve between proximal and distal parts; fore wing with usual three basal cells clearly enclosed by pigmented veins (costal, median and submedian cells); media vein (M) sclerotized and not reaching margin of wing ( Plate 52E, G ). Hindwing difficult to discern. Foreleg segments in following proportions: 8 (coxa): 3 (trochanter): 20 (femur): 10 (tibia): 7 (tarsal segment 1): 2 (tarsal segment 2): 2 (tarsal segment 3): 4 (tarsal segment 4): 9 (tarsal segment 5). Enlarged claw slightly shorter than tarsal segment 5 (7:9). Protrochanter short, not slender, not curved, without long proximal stalk. Shape of trochanter sinilar to those of Anteoninae . Segment 3 of protarsus produced into hook. Rudimentary claw apparently absent. Arolium much shorter than enlarged claw (1:7). Enlarged claw ( Plate 52H ) not spatulate, slightly curved, with distal apex slender and pointed, with inner side provided with one row of five bristles longer than greatest breadth of enlarged claw. Chela ( Plate 52H ) small and closed so that inner side of protarsal segment 5 hardly visible; distal apex of segment 5 with seven lamellae (ignored by Engel in his original description). According to original description, segment 5 of protarsus with one row of eight small lamellae (not seen by the authors) on inner side. Segment 5 of protarsus with proximal part slightly shorter than distal part. Midleg segments in following proportions: 10(coxa): 4 (trochanter): 24 (femur): 14 (tibia): 10 (tarsal segment 1): 5 (tarsal segment 2): 4 (tarsal segment 3): 3 (tarsal segment 4): 4 (tarsal segment 5). Hindleg segments in following proportions: 13 (coxa): 4 (trochanter): 25 (femur): 20 (tibia): 11 (tarsal segment 1): 5 (tarsal segment 2): 4.5 (tarsal segment 3): 3 (tarsal segment 4): 5 (tarsal segment 5). Petiole shape and length usual for Anteoninae . Shape, length and breadth of wings usual for Anteoninae . Shape and morphology of body usual for Anteoninae . Tibial spurs 1/1/2.

Male. Unknown.

Material examined. Type: holotype ♀ No. Bu-002: MYANMAR: specimen in Middle Cretaceous Burmese amber (about 100 mybp). Obtained from a mine situated in Northern Myanmar , Kachin State, Tanai Township , Hukawang Valley , SW of Tanai City ( NAMUK) .

Hosts. Unknown.

Distribution. Myanmar (Kachin).

Remarks. According to Grimaldi et al. (2002), on the basis of inclusions, Myanmar amber is from Lower Cenomanian. According to Cruikshank & Ko (2003) this amber is from Upper Albian, based on ammonite/pollen. Burmanteon Engel is very near the Neotropical genus Metanteon Olmi, 1984. According to Engel (2003), the differences between these two genera are the following:

Metanteon : distal abscissa Rs in fore wing terminating well before anterior wing margin (i.e. marginal cell apex broadly open); media vein (M) absent; proximal prominence absent on inner surface of enlarged claw.

Burmanteon : distal abscissa Rs in fore wing terminating near anterior wing margin (i.e. marginal cell apex narrowly open); media vein (M) present; proximal prominence present on inner surface of enlarged claw.

However, in Dryinidae the different length of distal abscissa Rs (=stigmal vein) is not a generic character; its value is only specific, as proved in many genera, such as Anteon Jurine, Dryinus Latreille, etc. In addition, the proximal prominence of the enlarged claw is considered absent in Metanteon , but it is present, although small (Plate 53). Probably the best distinctive character is the presence in Burmanteon of a strongly sclerotized Media vein (M). It is in fact fading in Metanteon . Another important character regards the length of the bristles present on enlarged claw inner margin: they are shorter than greatest breadth of enlarged claw in Metanteon (Plate 53), much longer in Burmanteon (Plate 52H).

Figures

PLATE 52. Burmanteon olmii, holotype. A. Lateral view. Length 1.6 mm. B. Head and antenna. C, D. Ventral view. E. Fore wing. F. Chela. G. Fore wing (basalmost portion of wing not illustrated) (from Engel, 2003). H. Chela (from Engel, 2003).