Pupina excisa Moellendorff , 1902

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina excisa Moellendorff, 1902

Figs 29H , 34E

Pupina (Tylotoechus) excisa Möllendorff, 1902a: 143. Type locality: Kelantan [Malaysia]. Laidlaw 1928: 34. Zilch 1957: 45, pl. 2, fig. 18.

Pupina excisa - Chan 1998a: 4, Ipoh, Perak. Chan 1998b: 2. Maassen 2001: 41. BEDO 2017: 90.

Type material examined.

Lectotype SMF 110778/1 (Figs 29H , 34E ) from Kelantan.

Diagnosis.

Shell ovate with higher spire; last whorl ca. three quarters of shell height. Apertural lip somewhat thickened but not expanded; apertural lip when observed from lateral view angled. Columellar tooth curved downward like an earlobe.

Differential diagnosis.

Pupina excisa can be distinguished from all other species in the P. arula species group from mainland Southeast Asia by an ovate shell shape with a higher spire, and an angled apertural lip when observed from lateral view. Pupina excisa is different from P. mouhoti in having a thicker, more prominent parietal tooth.

Distribution.

Kelantan and Perak States, Malaysia ( Maassen 2001).

Figures

Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 " forma minor ". Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J).

Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 " forma minor " from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B).