Greenwayodendron Verdc., Adansonia, ser . 2, 9 1. (1969)

Lissambou, Brandet-Junior, Hardy, Olivier J., Atteke, Christiane, Stevart, Tariq, Dauby, Gilles, Mbatchi, Bertrand, Sonke, Bonaventure & Couvreur, Thomas L. P., 2018, Taxonomic revision of the African genus Greenwayodendron (Annonaceae), PhytoKeys 114, pp. 55-93 : 55

Greenwayodendron Verdc., Adansonia, ser. 2, 9 1. (1969)

Polyalthia sect. Afropolyalthia Engler & Prantl., Leipzig,W. Engelmann.160. (1897)

Type species.

Greenwayodendron oliveri Engl.

Tree 2-45 m tall, d.b.h. 2-125 cm; stem cylindrical, bark smooth with large white stains, slice orange with a black ring, aromatic and rapidly turning brown. Young branches at first densely pubescent, later glabrous, trichomes 0.1-1.0 mm long; older branches sparsely pubescent to glabrous. Leaves entire, simple, alternate, astipulate; petiole 1.0-11.1 mm long, 0.8-3.3 mm in diameter, densely pubescent to glabrous, trichomes 0.1-1.2 mm long, indument brown; lamina 4.2-26.3 cm long, 2.0-9.6 cm wide, length: width ratio 1.5-4.0; elliptic to narrowly elliptic, base rounded or cuneate, apex acuminate, acute, apiculate, aristate or caudate, acumen 1-40 mm long, upper side sparsely pubescent to glabrous, lower side densely to sparsely pubescent; midrib upper side base densely pubescent to glabrous, lower side densely pubescent to glabrous, trichomes 0.1-1.2 mm long, indument tomentose; secondary veins 4-18 pairs, upper side sparsely pubescent to glabrous, lower side densely pubescent to glabrous, trichomes 0.1-1.0 mm long; tertiary veins upper side sparsely pubescent to glabrous, lower side sparsely pubescent to glabrous, irregularly prominent, slightly raised, distinct or indistinct above. Inflorescence axillary, 1-4 flowered rhipidium. Floral buds ellipsoid, 4-9 mm long, 2-5 mm in diameter, densely covered with short and long trichomes. Flowering pedicel 3.0-6.3 mm long, 0.5-2.2 mm in diameter, densely pubescent when young, becoming glabrous, trichomes 0.1-0.5 mm long; lower bract in lower half of pedicel, minute, sparsely pubescent; upper bract apical, just below the calyx, 0.9-5.2 mm in diameter, sparsely pubescent, trichomes 0.1-0.6 mm long. Sepals 1.3-5.0 mm long, 1.6-4.7 mm wide, length:width ratio 0.5-0.9 broadly ovate, imbricate, fused at the base, apex acuminate, base truncate, densely to sparsely pubescent outside, sparsely pubescent towards the centre inside, trichomes 0.1-0.5 mm long. Inner and outer petals subequal, 8.0-24.6 mm long, 1.3-3.5 mm wide, length:width ratio 0.5-0.9, narrowly ovate to elliptic, apex acuminate, base rounded, green maturing pale yellow, outside tomentose, trichomes 0.1-0.5 mm long, erect, inside sparsely pubescent to glabrous; glabrous part to 2-8 mm long. Male flowers: stamens 10-33, in several whorls, 1.2-4.2 mm long, 0.4-0.9 mm wide, tightly appressed; connective shield of stamens tongue-shaped, lobulated or flattened/discoid; hermaphrodite flowers: stamens 4-10 in a single whorl, appressed, 0.9-2.2 mm long and 0.3-0.8 mm wide, connective stamen tongue-shaped, lobulated or flattened; carpels 8-20, 0.7-2.1 mm long, 0.5-0.9 mm in diameter, length:width ratio 1.2-2.8 narrowly oblong, densely pubescent; ovules 1-2, oblong; stigmata ovoid, densely pubescent, trichomes 0.1-0.5 mm long. Fruiting pedicel 4.5-13.1 mm long, 1.0-3.5 mm in diameter, sparsely pubescent to glabrous, trichomes ca. 0.1-0.5 mm long; stipes 3.0-12 mm long and 1.0-3.2 mm in diameter, sparsely pubescent; monocarps 1-8, 2.5-21.1 mm in diameter, broadly ellipsoid to globose, sparsely pubescent to glabrous, green turning wine red at maturity; seeds 1-4 per monocarp, 3.0-15.4 mm in diameter, ellipsoid to globose, flattened on one side or hemi-ellipsoid when more than one seed per monocarp, surface covered by a white tegument; raphe impressed.