Quedius (Raphirus) ryvkini, Smetana & Shavrin, 2018

Smetana, Tephens & Shavrin, A., 2018, Contribution to the knowledge of the genus Quedius S, 1829 of Siberia and Russian Far East (Coleoptera: Staphylinidae: Staphylinini: Quediina), Linzer biologische Beiträge 50 (1), pp. 825-836 : 832-833

Quedius (Raphirus) ryvkini SMETANA nov.sp. ( Figs 12-16 )

T y p e l o c a l i t y. RUSSIA:PrimorieSchutzgebietSichote-Alin,Kordon Kabaniy.

T y p e m a t e r i a l. Holotype (♂), ROSSIA: " Russland: Primorie (S9) Schutzgebiet Sichote- Alin, Kordon Kabanyi 7.-13.VII.1998 leg. J. Sundukov / Coll. M. Schülke Berlin Ankauf Sundukow 1998", green plate. Deposited in MSC. Paratypes (2♂♂): "Primorskyi krai, Sikhote- Alinskyi zapovednik, Kabanyi kliuch. 45˚ 07’N 135˚ 52’E, 500 m. okonnye lovushki. 24- 27.04.2015. D.E. Shcherbakov" / " Quedius (Raphirus) sp.nov. 1 A. B. Ryvkin det. 2015". [Maritime Province, Sikhote-Alinskiy Nature Reserve, Kabanyi source, 45˚ 07’N 135˚ 52’E, 500 m. window traps 24- 27.04.2015. D. E. Shcherbakov]. Deposited in ZMM.

D i a g n o s i s. Species distinctive among the "larger" members of Raphirus with third antennal segment longer than the second, by the coloration of the body and appendages, in addition to the shape of the aedoeagus. Head black, pronotum piceous with each lateral margin rufotestaceous, elytra rufobrunneous to rufotestaceous, abdomen predominantly piceous-black to black; mandibles rufotestaceous, palpi, antennae and legs uniformly pale testaceous. Due to the coloration of the pronotum, the species resembles somewhat the specimens of the European Q. collaris ERICHSON, 1840 with lateral portions of pronotum paler, but it differs, in addition to the different aedoeagus, by the smaller, much less convex eyes.

D e s c r i p t i o n. Head black, pronotum piceous with each lateral margin rufotestaceous, elytra rufobrunneous to rufotestaceous, abdomen piceous-black to black with apical margins of tergites more or less narrowly paler. Mandibles rufotestaceous, palpi, antennae and legs uniformly pale testaceous. Head rounded, slightly wider than long (ratio 1.15); eyes large, moderately convex, tempora considerably shorter than length of eyes seen from above (ratio 0.31), no additional setiferous punctures between anterior frontal punctures; posterior frontal puncture separated from posteriomedial margin of eye by distance about equal to diameter of puncture; one basal puncture on each side; temporal puncture quite small, touching posterior margin of eye; surface of head with very fine and very dense microsculpture of mostly transverse waves. Antenna moderately long, slim, segment 3 longer than segment 2, segments 4 to 8 longer than wide, gradually becoming shorter, segment 9 and 10 about as long as wide, segment 11 about as long as two preceding segments combined. Pronotum visually as long as wide, but measured vaguely wider than long (ratio 1.08), widest at about midlength, narrowed anteriad, with lateral margins continuously arcuate with broadly rounded base, transversely convex, lateral portions not explanate; dorsal rows each with three punctures; sublateral rows each with one puncture situated before level of large lateral puncture; surface of pronotum with microsculpture similar to that on head, but even finer and denser, almost disappearing on lateral portions. Scutellum densely punctate/setose on apical half, surface with rudimentary microsculpture. Elytra moderately long, at base slightly narrower than pronotum at widest point, vaguely dilated posteriad; at suture slightly shorter (ratio 0.89) and at sides about as long as pronotum at midline; punctation very fine and very dense, transverse interspaces between punctures mostly about as large as diameters of punctures; surface between punctures with microscopic micropunctulation, elytra therefore appearing slightly opaque; pubescence testaceous. Wings fully developed. Abdomen with tergite 2 (in front of first fully visible tergite) only with fine setiferous punctures at apical margin, otherwise impunctate; tergite 7 (fifth visible) with fine apical seam of palisade setae; punctation of tergites similar to that on elytra, becoming somewhat sparser toward apical margin of each tergite and in general toward apex of abdomen; pubescence piceous; surface between punctures with excessively fine microsculpture of transverse striae.

Male. First four segments of front tarsus moderately dilated, sub-bilobed, each with tenent setae ventrally; segment 2 about as wide as apex of tibia; segment 4 narrower than preceding segments. Sternite 8 with three strong setae at each side, with narrow, obtusely triangular medioapical emargination, narrow triangular area before emargination flattened and smooth ( Fig. 12 ). Genital segment with tergite 10 evenly narrowed toward arcuate apex, with several long setae at apex and numerous short setae in front of them ( Fig. 13 ); sternite 9 with small, wide basal portion, apical portion subarcuate at apex, with stronger apical seta in each lateroapical corner, otherwise finely and densely setose ( Fig. 14 ). Aedoeagus ( Figs 15, 16 ) narrow, elongate, median lobe slightly, widely attenuated at about apical third, anteriorly narrowed into apical portion with narrowly arcuate apex; paramere large, elongate, covering median lobe except for narrow lateral portions, anteriorly narrowed into subacute apex, apex not quite touching apex of median lobe; four apical setae, median pair markedly longer than lateral pair and two long setae at each lateral margin close to apex; underside with quite numerous sensory peg setae, covering entire apical third of underside.

Female. unknown.

Length 7.5-8.0 mm.

E t y m o l o g y. Patronymic, the species was named in honor of our colleague Alexandr B. Ryvkin, Moscow, Russia, who correctly recognized its taxonomic status and gratuitously made the specimens available to senior author.

G e o g r a p h i c a l d i s t r i b u t i o n. Quedius ryvkini is at present known only from the type locality in Sikhote-Alin range in the Maritime Province of the Russian Far East.

B i o n o m i c s. Two of the known three s pecimens were taken from window traps, but it is not known in what habitat the traps were set.

C o m m e n t. The setation of sternite 9 of male genital segment was damaged, Fig. 14 shows the preserved setation.

Figures

Figs 12-21: 12-16. Quedius ryvkini nov.sp. (12) apical portion of male sternite 8; (13) tergite 10 of male genital segment; (14) sternite 9 of male genital segment; (15) aedoeagus, ventral view; (16) apical portion of underside of paramere with sensory peg setae. 17-21. Quedius aedilis nov.sp. (17) apical portion of male sternite 8; (18) tergite 10 of male genital segment; (19) sternite 9 of male genital segment; (20) aedoeagus, ventral view; (21) apical portion of underside of paramere with sensory peg setae.