Wiedemannia nebulosa Ivkovic & Sinclair

Ivkovic, Marija, Cevid, Josipa, Horvat, Bogdan & Sinclair, Bradley J., 2017, Aquatic dance flies (Diptera, Empididae, Clinocerinae and Hemerodromiinae) of Greece: species richness, distribution and description of five new species, ZooKeys 724, pp. 53-100 : 58-59

Wiedemannia nebulosa Ivkovic & Sinclair sp. n. Figs 1, 9

Type locality.

Greece: Thrace, north of Dipotama, 41°24'24"N, 24°37'19"E, 1400 m.

Type material.

Holotype ♂, labelled: "GREECE: Thrace/ N of Dipotama/ 41°24'24"N, 24°37'19"E / 23.v.1994; 1400 m/ leg. B. Horvat, I. Sivec"; "HOLOTYPE/ Wiedemannia / nebulosa / Ivković & Sinclair" (CNC, dried from alcohol). Paratypes: same data as holotype (1 ♂, 1 ♀, CNC, dried from alcohol).

Diagnosis.

This species of Wiedemannia is distinguished by the faint clouding about crossveins and base of radial fork, shape of the clasping cercus and position of distiphallus on the phallic shaft.

Description.

Male. Body length 3.8-4.5 mm, wing length 5.2-5.3 mm (colouration slightly bleached by prolonged storage in alcohol). Head dark with brown frons and vertex, remainder of head with blue pruinescence; head higher than long; gena narrow, one-quarter height of eye. Frons short, broader than face. Face wide, with distinct carina on lower margin, bare, lacking setae. One pair of long ocellar setae and one pair of vertical setae; 6-7 distinct upper postoculars; lower postocular setae finer and merging with longer setae on middle and lower occiput; a few small setulae present on vertex and in ocellar area. Antenna brown; postpedicel and stylus minutely pubescent; scape longer than pedicel, with setulae dorsally; pedicel with complete circlet of apical setae; postpedicel apically pointed; stylus twice length of postpedicel.

Scutum dark brown with pair of faint black vittae between dorsocentral row and acrostichals and bluish stripe medially; prescutellar depression with blue pruinescence. Pleura clothed with blue pruinescence. Mesonotum with 5 pairs of dorsocentral setae without short setulae interspersed. Acrostichal setae short and fine, biserial, extending onto prescutellar depression; 1 strong postpronotal seta; 2 notopleural setae and several short setae; 1 presutural supra-alar seta and several small anterior setulae; 1 postalar seta. Antepronotum with 1 pair of strong setae. Proepisternum with some fine setulae. Katepisternum without setulae. Laterotergite with fine, pale setae. One pair of strong marginal scutellar setae; disc bare.

Wing membrane infuscate with darkening at apex of cell dm, radial fork and r-m crossvein; veins darker; 1 short basal costal seta ending before humeral crossvein. Cell dm produced anteroapically. M1 and M2 originating separately from cell dm. CuA+CuP in form of short streak. Pterostigma broad and elongate, very distinct. Squama with setulae. Halter yellowish brown.

Legs mostly brown; fore femur with 2-3 strong anterior setae on apical quarter; uniformly covered with rows of small dark setulae. All coxae with longer setae anteriorly; fore coxa with 1-2 erect setae. Fore and mid femora ventrally with some longer setulae on proximal half.

Abdomen concolourous with thorax, covered in short setae. Pruinescence darker on tergites than sternites. Terminalia (Fig. 9): hypandrium subequal in length with epandrium, with 5 pairs of setae. Epandrium irregularly subquadrate, with several stouter and longer setae (shown by enlarged sockets) in addition to normal setae ventrally and laterally; surstylus short, digitiform with rounded apex; subepandrial sclerite projecting slightly beyond epandrium near surstylus. Clasping cercus pale brown, broad, gradually tapered to rounded apex; inner posterior margin with long peg-like setae. Phallus more or less linear, slender; distiphallus without swelling at mid-length; distiphallus with serrate membranous margin, extending onto shaft.

Female. Similar to male. Terminalia: cercus short ovate and minutely pilose.

Etymology.

The species name is derived from the Latin nebulosus (misty, cloudy, dark), in reference to the clouding about the crossveins.

Remarks.

Wiedemannia nebulosa sp. n. is known only from the type locality in Greece. On the basis of the shape of the clasping cercus, this new species is similar to W. carpathica Vaillant, 1967 (eastern Carpathians), W. pyrenaica Vaillant, 1967 (Pyrenees) and perhaps W. wachtli (Mik, 1880).