Acylomus Sharp, 1888

Gimmel, Matthew L., 2013, <strong> Genus-level revision of the family Phalacridae (Coleoptera: Cucujoidea) </ strong>, Zootaxa 3605 (1), pp. 1-147 : 28-35

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https://doi.org/ 10.11646/zootaxa.3605.1.1

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scientific name

Acylomus Sharp, 1888
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4. Acylomus Sharp, 1888

( Figs. 2l View FIGURE 2 ; 3a View FIGURE 3 ; 6–8 View FIGURE 6 View FIGURE 7 View FIGURE 8 ; 37g –i View FIGURE 37 )

Acylomus Sharp 1888: 256 . Type species: Acylomus aciculatus Sharp 1889 , fixed by monotypy.

Liophalacrus Sharp 1888: 255 . Type species: Liophalacrus bicolor Sharp 1888 , fixed by subsequent designation. Syn. nov.

Coelocoelius Guillebeau 1893 a: 290 . Type species: Coelocoelius simoni Guillebeau 1893 , fixed by monotypy. [synonymized with Acylomus Sharp by Champion (1924 c: 244)]

Ganyrus Guillebeau 1894 a: 280 . Type species: Ganyrus rubellus Guillebeau 1894 , fixed by original designation. Syn. nov.

Podocesus Guillebeau 1894 a: 281 . Type species: Eustilbus semirufus Guillebeau 1893 , fixed by original designation. Syn. nov.

Tinodemus Guillebeau 1894 a: 282 . Type species: Tinodemus grouvellei Guillebeau 1894 , fixed by original designation. Syn. nov.

Dolerus Guillebeau 1894 a: 282 . Type species: Dolerus limbatus Guillebeau 1894 , fixed by original designation.

Ledorus Guillebeau 1895 : xxvii. Type species: Dolerus limbatus Guillebeau 1894 , fixed by objective synonymy with Dolerus Guillebeau. [replacement name for Dolerus Guillebeau, 1894 ] [synonymized with Podocesus Guillebeau by Švec (2003: 117)] Syn. nov.

Astenulus Guillebeau 1896: 299 . Type species: Astenulus micropus Guillebeau 1896 , fixed by monotypy. [synonymized with Tinodemus Guillebeau by Švec (2002 b: 220)]. Syn. nov.

Afronyrus Švec 2006: 106 . Type species: Afronyrus snizeki Švec 2006 , fixed by original designation. Syn. nov.

Type material. Acylomus aciculatus Sharp : 18 syntypes found in BMNH, one dissected male, point mounted, card containing left protibia/tarsus, right maxillary palp, and right antenna, tegmen and median lobe in glycerol-filled capsule, here designated as a lectotype to stabilize the species and generic name, “Sp. figured // Rio Hondo, \ B. Honduras. \ Blancaneau. // Acylomus aciculatus, Ch. [handwritten] // B.C.A.,Col., II,(1). \ Acylomus aciculatus . // LECTOTYPE \ Acylomus \ aciculatus Sharp [binomial handwritten] \ W.E. Steiner, Jr. [red label, designation not published, turned over] // LECTOTYPE \ Acylomus \ aciculatus Sharp \ des. M.L. Gimmel 2011 [red label]” ( BMNH). Paralectotypes: 17 ( BMNH), with label attached “ PARALECTOTYPE \ Acylomus \ aciculatus Sharp \ det. M.L. Gimmel 2011 [yellow label]”.

Liophalacrus bicolor Sharp : eight syntypes found in BMNH, the card-mounted specimen with “Type” handwritten by David Sharp selected as the lectotype to stabilize the species and generic name, “ Liophalacrus \ bicolor. \ Type D.S. \ Bugaba Champion. [handwritten on specimen card] // Type [red-bordered disc] // Bugaba, Panama. \ Champion // Sharp Coll. \ 1905.–313. // LECTOTYPE \ Liophalacrus \ bicolor Sharp \ des. M.L. Gimmel 2011 [red label]” ( BMNH). Paralectotypes: 7 card-mounted specimens, with label attached “ PARALECTOTYPE \ Liophalacrus \ bicolor Sharp \ des. M.L. Gimmel 2011 [yellow label]” ( BMNH).

Coelocoelius simoni Guillebeau : two syntypes, card mounted, “San Esteban \ E. Simon III.88” ( MNHN). Only two of the supposed four syntypes were found.

Ganyrus rubellus Guillebeau : holotype, female, “Abyss. \ Raffray [blue label] // 167 // Grouvelle [handwritten] // [handwritten label, illegible] // HOLOTYPE ♀ \ Ganyrus \ rubellus Guillebeau \ det. M.L. Gimmel 2009 [red label]” ( MNHN), point mounted, genitalia in DMHF.

Eustilbus semirufus Guillebeau : holotype, male, card mounted, “Caracas \ 1 88 E S // Simon // TYPE // Museum Paris \ Coll. Générale // Lectotypus \ PODOCESUS SEMIRUFUS Guill. 1894 \ Z. Svec des. 1999 // GENITALIA IN WATER SOLUBLE MEDIUM — DMHF // semirufus Guilb. ” ( MNHN). The lectotype designation is in error (the species was described from “1 exemplaire”).

Tinodemus grouvellei Guillebeau : lectotype, male, card mounted, genitalia dissected, “ Michigan // Grouvelle // Museum Paris \ Coll. Générale // TYPE // Lectotypus TINODEMUS GROUVELLEI Guillebeau 1894 \ Z. Svec des. 1999 // GENITALIA IN DMHF — WATER SOLUBLE MED. // Grouvellei Guilb. // Acylomus \ ergoti Casey \ det. M. Gimmel 2008” ( MNHN).

Dolerus limbatus Guillebeau : holotype, female, card mounted, “ Grouvelle // Colombie // Museum Paris \ Coll. Générale // TYPE // Lectotypus DOLERUS LIMBATUS Guillebeau 1894 \ Z. Svec des. 1999 // ANTENNA IN DMHF — WATER SOLUBLE MEDIUM // limbatus Guilb. // Ledorus // Dolerus // limbatus Guilb. ” ( MNHN). The lectotype designation is in error (the species was described from “1 exempl.”).

Astenulus micropus Guillebeau : holotype, male, genitalia dissected, “Alluaud // Diego Suarez // Museum Paris \ Coll. Générale // HOLOTYPE ” ( MNHN) .

Afronyrus snizeki Švec : type not accessible.

Diagnosis. May be recognized by the divergent metaventral postcoxal lines which may be arcuate or angulate, single elytral sutural stria, metatarsomere I shorter than II with joint between them more or less rigid, prosternal process angulate when viewed laterally and usually with row or pair of stiff setae at apex, ventral lobe of the eye not expanded posteriorly, mandible without a ventral ridge, and the tegmen with parameres hinged to basal piece.

Description. Very small to large, total length 1.3–3.5 mm. Dorsal color usually dark reddish-brown to piceous, sometimes with apex of elytra paler, or with pale maculations on disc ( Fig. 37g –i View FIGURE 37 ). Tibial spur formula 2-2-2, tarsal formula 5-5- 5 in females, 5-5- 4 in males.

Head. Not constricted behind eyes ( Fig. 2l View FIGURE 2 ). Eyes small to medium-sized; facets flat; interfacetal setae absent; strongly emarginate to straight medially; often with sharp posterior emargination; periocular groove present; with transverse setose groove ventrally behind eye. Frontoclypeus emarginate above antennal insertion; clypeal apex arcuate-truncate. Antennal club 3-segmented, club symmetrical, antennomere XI not constricted ( Figs. 6b View FIGURE 6 , 7b View FIGURE 7 , 8b View FIGURE 8 ). Mandible ( Figs. 6a View FIGURE 6 , 7a View FIGURE 7 , 8a View FIGURE 8 ) with apex bidentate; with retinaculum; mandible without ventral ridge. Maxillary palpomere IV fusiform, inner edge swollen medially; galea short, rounded; lacinia with two stout spines, often with associated tuft of setae. Mentum with sides divergent toward apex; labial palpomere III fusiform. Labrum with apical margin arcuate to emarginate. Gular sutures short, barely evident, rarely long.

Thorax. Pronotum with or without obvious microsetae; with weakly to moderately developed scutellar lobe. Prosternum anteriorly with marginal setae distributed in two patches, setae normal; procoxal cavity with anterolateral notchlike extension; prosternal process angulate in lateral view, not conspicuously setose preapically, usually with row or pair of spinelike setae at apex ( Fig. 3a View FIGURE 3 ). Protrochanter without setae; protibia without ctenidium on kickface ( Figs. 6c View FIGURE 6 , 7c View FIGURE 7 , 8c View FIGURE 8 ). Scutellar shield small. Elytron usually without spectral iridescence, rarely present; one sutural stria present; discal striae absent or barely suggested; without or with weak transverse strigae; lateral margin usually with row of tiny, sawtooth-like setae. Mesoventral plate ( Figs. 6f View FIGURE 6 , 7e View FIGURE 7 , 8f View FIGURE 8 ) notched anteriorly, extending posteriorly to metaventrite, dividing mesoventral disc in two, not forming or forming weak procoxal rests; mesanepisternum with incomplete transverse carina; mesocoxal cavities widely separate, separated by more than half width of a coxal cavity. Mesotarsomere III bilobed or not. Metaventral process ( Figs. 6f View FIGURE 6 , 7e View FIGURE 7 , 8f View FIGURE 8 ) not extending anteriorly beyond anterior level of mesocoxae; metaventral postcoxal lines diverging from mesocoxal cavity margin, arcuate and smoothly rounded to acuminately pointed, branches always connected, never with a spur; discrimen short, extending less than halfway to anterior margin of metaventral process; metendosternite ( Figs. 6g View FIGURE 6 , 7f View FIGURE 7 , 8g View FIGURE 8 ) with anterior tendons moderately separated, ventral process intersecting ventral longitudinal flange behind anterior margin. Anterior margin of metacoxa with emargination sublaterally; metacoxal plate with transverse line present or absent; metatibial foreface with apical ctenidium roughly perpendicular overall to long axis of tibia; spurs cylindrical, longest spur subequal in length to or longer than width of tibial apex, spurs ( Fig. 6d View FIGURE 6 ) and tibial apex sometimes modified in males; metatarsus slender, metatarsomere I distinctly shorter than metatarsomere II, joint between I and II rigid ( Figs. 6d View FIGURE 6 , 7d View FIGURE 7 , 8d View FIGURE 8 ). Hind wing ( Figs. 6e View FIGURE 6 , 8e View FIGURE 8 ) with distinct, ovate anal lobe; leading edge with complete row of long setae at level of RA +ScP; AA 3+4 not apparent; cubitoanal system often branched apically; CuA 2 and MP 3+4 usually without distal remnants; r4 present or absent; flecks present in apical field just distal to rp-mp2; long to short transverse proximal sclerite and additional strong or moderate, irregular sclerite present just distal to end of radial bar.

Abdomen. Abdominal ventrite I without paired lines or calli; spiracles apparently absent from segment VII. Male with aedeagus upright in repose; tegmen ( Figs. 6h View FIGURE 6 , 7g View FIGURE 7 , 8h View FIGURE 8 ) with symmetrical anterior margin, parameres hinged to basal piece, parameres with or without medial longitudinal division; penis ( Figs. 6i View FIGURE 6 , 7h View FIGURE 7 , 8i View FIGURE 8 ) variable, with endophallic spicules, often with large sclerites, apex often pointed or tripartite; spiculum gastrale V-shaped, with arms free. Female ovipositor weakly sclerotized, palpiform.

Immature stages. The larva and pupa of Acylomus pugetanus Casey were described and illustrated by Steiner and Singh (1987).

Bionomics. Most members of this genus, at least in the Nearctic region, appear to be generalist ascomycete fungus grazers on dead vegetation as adults and larvae. Dead hanging leaf clusters, a habitat described in Steiner (1984), seems to be especially favored by a few eastern Nearctic species. I have observed one species, A. elongatulus , in large numbers on grass tussocks in Louisiana. Another species, A. pugetanus , develops within the sclerotia of Claviceps species (ergot) on grasses in northern North America (see Steiner and Singh 1987 for details). Most members are attracted to lights at night, often in large numbers.

Distribution and diversity. A widely distributed genus, well represented in most tropical and subtropical regions and in eastern North America, but absent over much of the Palearctic region. I have examined at least three species of this genus from Australia, with most specimens from the northern half of the continent; whether these are undescribed has not been determined. Although many synonyms apparently exist in the genus, a great many species are undescribed. Upon revision this will likely become the most species-rich phalacrid genus, should it prove to be monophyletic.

Included species (94):

Acylomus abjectus Casey, 1916 ( Distribution : United States) (type!)

Acylomus aciculatus Sharp, 1889 ( Distribution : Central America) (type!)

Acylomus acuminatus ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: Tanzania)

Acylomus acutangulus (Kirsch, 1873) , comb. nov. ( Phalacrus ) ( Distribution : Peru) (type!)

Acylomus ambagiosus ( Lyubarsky, 2003) , comb. nov. ( Stilbus ) (Distribution: Nepal)

Acylomus apicalis ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: Kenya)

Acylomus atomarius ( Sharp, 1888) , comb. nov. ( Olibrus ) ( Distribution : Panama) (type!)

Acylomus bicolor ( Sharp, 1888) , comb. nov. ( Liophalacrus ) ( Distribution : Panama) (type!)

Acylomus bicoloratus Gimmel , nom. nov. [for Tinodemus bicolor Švec, 2002 , junior secondary homonym of Acylomus bicolor ( Sharp, 1888) ] (Distribution: Tanzania)

Acylomus bifurcus ( Švec, 1992) , comb. nov. ( Tinodemus ) ( Distribution : Japan) (type!)

Acylomus borealis (Guillebeau, 1894) (Distribution: Canada) (type!)

Acylomus calcaratus Casey, 1890 (Distribution: Bahamas, Bermuda, United States) (type!)

Acylomus capriviensis ( Lyubarsky, 1998) , comb. nov. ( Podocesus ) (Distribution: southern Africa)

Acylomus carbonarius Casey, 1916 ( Distribution : United States) (type!)

Acylomus championi (Hetschko, 1929) , comb. nov. ( Tinodemus ) ( Distribution : Namibia, South Africa) (type!)

Acylomus chinensis ( Švec, 1992) , comb. nov. ( Tinodemus ) ( Distribution : China) (type!)

Acylomus claviger (Champion, 1925) , comb. nov. ( Tinodemus ) (Distribution: subsaharan Africa) (type!)

Acylomus confusus Casey, 1916 ( Distribution : United States) (type!)

Acylomus confusus ( Švec, 1992) , comb. nov. ( Tinodemus ) ( Distribution : Japan) (type!) [junior secondary homonym not replaced since the older name is probably a synonym]

Acylomus cubensis Casey, 1916 ( Distribution : Cuba) (type!)

Acylomus curvolineatus (Champion, 1924) , comb. nov. ( Stilbus ) ( Distribution : Oriental region) (type!) [see note on synonymy below]

Acylomus darwinii (Waterhouse, 1877) (Distribution: Ecuador) (type!)

Acylomus detractus Casey, 1916 ( Distribution : Cuba) (type!)

Acylomus digestus Casey, 1916 ( Distribution : United States) (type!)

Acylomus distinctus ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: southern Africa)

Acylomus ellipticus Casey, 1916 ( Distribution : United States) (type!)

Acylomus elongatulus (Casey, 1890) (Distribution: United States) (type!)

Acylomus ergoti Casey, 1890 ( Distribution : United States) (type!)

Acylomus eximius Casey, 1916 ( Distribution : United States) (type!)

Acylomus extricatus Casey, 1890 ( Distribution : United States) (type!)

Acylomus flaviceps (Guillebeau, 1894) , comb. nov. ( Tinodemus ) ( Distribution : Colombia) (type!)

Acylomus fortis Champion, 1925 (Distribution: Brazil) (type!) Acylomus grouvellei (Guillebeau, 1894) , comb. nov. ( Stilboides ) (Distribution: Brazil, Cuba) (type!) Acylomus humilis Casey, 1916 (Distribution: United States) (type!) Acylomus insularis (Guillebeau, 1894) (Distribution: Martinique) (type!) Acylomus integer Casey, 1916 (Distribution: United States) (type!) Acylomus interpositus ( Švec, 1992) , comb. nov. ( Podocesus ) (Distribution: Japan) (type!) Acylomus latisternus (Guillebeau, 1894) (Distribution:? Haiti) (type!) Acylomus libidinosus ( Lyubarsky, 2003) , comb. nov. ( Stilbus ) (Distribution: Vietnam) Acylomus limbatus (Guillebeau, 1894) , comb. nov. ( Podocesus ) (Distribution: Colombia) (type!) Acylomus lyubarskyi Gimmel , nom. nov. [for Olibrus capriviensis Lyubarsky, 1998 , junior secondary homonym of Acylomus capriviensis ( Lyubarsky, 1998) ] (Distribution: Namibia) Acylomus maruskae ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: Kenya, Tanzania, Uganda) Acylomus mesomelas (Champion, 1925) , comb. nov. ( Tinodemus ) (Distribution: South Africa, Tanzania,

Zimbabwe) (type!) Acylomus mexicanus ( Sharp, 1888) , comb. nov. ( Olibrus ) (Distribution: Belize, Guatemala, Mexico) (type!) Acylomus micaceus Casey, 1916 (Distribution: Mexico) (type!) Acylomus micropus ( Guillebeau, 1896) , comb. nov. ( Tinodemus ) (Distribution: Madagascar, Réunion) (type!) Acylomus mifsudi (Švec, 2000) , comb. nov. ( Tinodemus ) (Distribution: Malta) Acylomus morosus Casey, 1916 (Distribution: United States) (type!) Acylomus nebulosus Casey, 1890 (Distribution: United States) (type!) Acylomus neglectus ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: Guinea, Zambia) Acylomus oblongus (Guillebeau, 1894) , comb. nov. ( Tinodemus ) (Distribution: Brazil) (type!) Acylomus obsoletus ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: Kenya) Acylomus obtusus ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: South Africa) Acylomus orientalis Gimmel , nom. nov. [for Stilbus similis Švec, 1992 , junior secondary homonym of

Acylomus similis ( Scott, 1922) ] (Distribution: China, Japan) (type!) Acylomus ornatus (Guillebeau, 1894) , comb. nov. ( Tinodemus ) (Distribution: Mexico) (type!) Acylomus ovalis ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: Tanzania, Uganda) Acylomus ovulatus Casey, 1916 (Distribution: United States) (type!) Acylomus partitus ( Sharp, 1888) , comb. nov. ( Olibrus ) (Distribution: Guatemala) (type!) Acylomus parvulus ( Boheman, 1858) , comb. nov. ( Olibrus ) (Distribution: Peru) Acylomus piceus Casey, 1890 (Distribution: United States) (type!) Acylomus pictus (Horn, 1896) , comb. nov. ( Litolibrus ) (Distribution: Mexico) (type!) Acylomus polygramma (Flach, 1888) , comb. nov. ( Tinodemus ) (Distribution: Mediterranean region) Acylomus porrectus ( Sharp, 1888) , comb. nov. ( Olibrus ) (Distribution: Central America) (type!) Acylomus pugetanus Casey, 1916 (Distribution: Canada, United States) (type!) Acylomus pumilus (Guillebeau, 1894) , comb. nov. ( Ganyrus ) (Distribution: Indonesia) (type!) Acylomus quadrispinosus Casey, 1916 (Distribution: Cuba) (type!) Acylomus reticulatus (Guillebeau, 1894) , comb. nov. ( Ganyrus ) (Distribution: Indonesia) (type!) Acylomus rotundus ( Sharp, 1888) , comb. nov. ( Liophalacrus ) (Distribution: Panama) (type!) Acylomus rubellus (Guillebeau, 1894) , comb. nov. ( Ganyrus ) (Distribution: Ethiopia) (type!) Acylomus rubicundus (Champion, 1925) , comb. nov. ( Tinodemus ) (Distribution: Namibia, Zimbabwe) (type!) Acylomus ruficornis ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: Kenya) Acylomus rufopunctatus ( Lyubarsky, 1998) , comb. nov. ( Podocesus ) (Distribution: Namibia, South Africa,

Tanzania) Acylomus sanderi ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: subsaharan Africa) Acylomus sculpturatus ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: Guinea) Acylomus secundus ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: subsaharan Africa) Acylomus semirufus (Guillebeau, 1893) , comb. nov. ( Podocesus ) (Distribution: Venezuela) (type!) Acylomus similis ( Scott, 1922) , comb. nov. ( Nesiotus ) (Distribution: Seychelles) (type!) Acylomus simoni (Guillebeau, 1893) (Distribution: Venezuela) (type!) Acylomus snizeki ( Švec, 2002) , comb. nov. ( Tinodemus ) (Distribution: Guinea, Uganda) Acylomus socialis Casey, 1916 (Distribution: United States) (type!)

Acylomus stilboides (Guillebeau, 1894) (Distribution: Brazil) (type!)

Acylomus strigillatus (Guillebeau, 1894) , comb. nov. ( Ganyrus ) ( Distribution : Mexico) (type!)

Acylomus subhemisphaericus (Guillebeau, 1894) (Distribution: Brazil) (type!)

Acylomus submaculatus ( Sharp, 1888) , comb. nov. ( Olibrus ) (Distribution: Central America) (type!)

Acylomus substrigosus ( Sharp, 1888) , comb. nov. ( Olibrus ) ( Distribution : Guatemala) (type!)

Acylomus sveci Gimmel , nom. nov. [for Tinodemus reticulatus Švec, 2002 , junior secondary homonym of Acylomus reticulatus (Guillebeau, 1894) ] (Distribution: South Africa, Tanzania)

Acylomus teapensis ( Sharp, 1888) , comb. nov. ( Olibrus ) ( Distribution : Mexico) (type!)

Acylomus texanus Casey, 1916 ( Distribution : United States) (type!)

Acylomus tropicus ( Scott, 1922) , comb. nov. ( Tinodemus ) ( Distribution : Réunion, Seychelles) (type!)

Acylomus vacivus Casey, 1916 ( Distribution : United States) (type!)

Acylomus versicolor (Kirsch, 1873) , comb. nov. ( Olibrus ) ( Distribution : Peru) (type!)

Acylomus vicinus (Guillebeau, 1894) (Distribution: Brazil) (type!)

Acylomus vividus Casey, 1916 ( Distribution : United States) (type!)

Acylomus zdeneki Gimmel , nom. nov. [for Afronyrus snizeki Švec, 2006 , junior secondary homonym of Acylomus snizeki ( Švec, 2002) ] (Distribution: Kenya)

Discussion. Based on the original description ( Boheman 1858), Olibrus parvulus cannot belong to Olibrus , for it has only a single sutural stria and the type is from Peru. It is likely a member of Acylomus , and I have tentatively transferred it to this genus.

Into this genus I have placed all New World Stilbus -group members with hinged parameres and without the characters of Xanthocomus . This includes the type species of Podocesus , P. semirufus Guillebeau (illustrated in Švec 2003: figs. 37–44), the type species of Ledorus , Dolerus limbatus Guillebeau (illustrated in Švec 2003: figs. 49, 50), and the type species of Tinodemus , T. grouvellei Guillebeau (illustrated in Švec 2002: figs. 9–16). In fact, T. grouvellei , described from “ Michigan,” is identical in aedeagal characteristics to a previously described form, Acylomus ergoti Casey. I therefore consider these two to be synonyms:

Acylomus ergoti Casey 1890 = Tinodemus grouvellei Guillebeau 1894 , syn. nov.

Coelocoelius was synonymized with Acylomus by Champion (1924 c). I have examined the type of the type species, C. simoni , and concur with this assessment.

The type of the type species of Ganyrus , G. rubellus Guillebeau ( Ethiopia) , falls well within my concept of Acylomus , and therefore I am synonymizing the two. Externally it is similar to Acylomus sanderi (Švec) . I have seen the types of the three other species that were described in Ganyrus : G. strigillatus Guillebeau ( Mexico) , which obviously belongs in Acylomus ; G. pumilus Guillebeau and G. reticulatus Guillebeau (both Sumatra), whose generic assignment to Acylomus is tentative given the condition of the types, whose ventral surfaces were obscured. Examination of the syntype series of Nesiotus similis Scott (1922: 239) has revealed that this species is more properly placed in Acylomus , based on the normally-proportioned antennal club and tarsal structure. Its metaventral postcoxal lines are arcuate, excluding the species from Stilbus Seidlitz.

Dissection of the lectotype (BMNH), here designated (complete label data: “Sarda, \ Bengal \ F. W. C. // G.C. Champion. \ Brit. Mus. \ 1925–42. // Stilbus \ curvolineatus, \ Champ. // E.M.M. 1924. \ det. G.C.C. // SYN- \ TYPE [blue-bordered disc] // LECTOTYPE ♂ \ Stilbus \ curvolineatus Champion \ des. M.L. Gimmel 2010 [red label]”), of Stilbus curvolineatus Champion ( India) reveals an aedeagus much like that illustrated for both Tinodemus meridianus (Švec) ( Afghanistan, Japan) and Olibrus stuporatus Lyubarsky (Java, Nepal) along with their original descriptions. I am considering these three names as synonymous, and A. curvolineatus assumes priority. The lectotype is designated to prevent future doubts about the identity of the species. The (one) paralectotype is female.

Acylomus curvolineatus (Champion 1924) = Tinodemus meridianus ( Švec 1992) = Olibrus stuporatus

Lyubarsky 1994, syn. nov.

I have been unable to examine the type of Afronyrus snizeki Švec located in Švec’s private collection, but based on the description and illustrations it falls within my broadened concept of Acylomus , based on the parameres hinged to the basal piece and modified tibial spurs of the male. Therefore I consider Afronyrus a junior synonym of Acylomus . This placement (and others in this genus) may be revised when detailed studies of the species and species groups of this complex genus are undertaken.

MNHN

Museum National d'Histoire Naturelle

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Phalacridae

Loc

Acylomus Sharp, 1888

Gimmel, Matthew L. 2013
2013
Loc

Afronyrus Švec 2006: 106

Svec, Z. 2006: 106
2006
Loc

Ledorus

Svec, Z. 2003: 117
2003
Loc

Astenulus

Guillebeau, F. 1896: 299
1896
Loc

Ganyrus

Guillebeau, F. 1894: 280
1894
Loc

Podocesus

Guillebeau, F. 1894: 281
1894
Loc

Tinodemus

Guillebeau, F. 1894: 282
1894
Loc

Dolerus

Guillebeau, F. 1894: 282
1894
Loc

Coelocoelius

Champion, G. C. 1924: 244
Guillebeau, F. 1893: 290
1893
Loc

Acylomus

Sharp, D. 1888: 256
1888
Loc

Liophalacrus

Sharp, D. 1888: 255
1888
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