Fissocantharis securiclata Y. Yang & X. Yang

Yang, Yuxia, Qi, Yaqing & Yang, Xingke, 2018, Four new species of Fissocantharis Pic, 1921 (Coleoptera, Cantharidae) from China, ZooKeys 738, pp. 97-115 : 101

Fissocantharis securiclata Y. Yang & X. Yang sp. n. Figs 1A, 2 A–C, 4A, 5A

Type material.

Holotype ♂ (MHBU): CHINA, Anhui, Jixi, Qingliangfeng, 2-5.VI.2013, leg. J.S. Xu & C.X. Yuan. Paratypes: CHINA: 1♀ (MHBU): same data as holotype; Zhejiang: 1♀ (IZAS): "Tienmu Shan, 10.6.1936, O. Piel coll."; 1♂ (IZAS): "Tienmu Shan, 6.6.1936, O. Piel coll."; 1♂ (CoAP) "CHINA: Zhejiang [CH07-39], Hangzhou Pref., Tianmu Shan, 40 km WNW Linan, water reservoir, 30°20'56"N, 119°18'42"E, 300 m, plant refuse, litter from rock edges, 17.VI.2007, leg. A. Pütz”.

Distribution.

China (Anhui, Zhejiang).

Description.

Male (Fig. 1A). Head orange, darkened at both sides of vertex, mouthparts orange, darkened at mandibular apices, terminal maxillary and labial palpomeres and antennae orange, darkened dorsally at antennomeres III–IX, pronotum, scutellum and elytra black, legs orange, darkened at tarsi, ventral surface of body black, yellow at posterior and sides of abdominal sternites. Body densely covered with short recumbent gray pubescence.

Head subquadrate, narrowed posteriorly behind eyes; eyes moderately projecting, head width across eyes greater than that of anterior edge of pronotum; apical maxillary palpomeres nearly long-triangular, widest at apical two-fifths, acute at apices; antennae extending to elytral midlength, antennomeres II about twice as long as wide, III obliquely widened apically, truncate at apical angles, about twice as long as wide, IV–IX securiform, abruptly widened apically and rounded at apical angles, IV and IX slightly longer than wide, V–VIII nearly as long as wide, X and XI nearly parallel-sided, XI pointed at apices and about 1.5 times as long as X.

Pronotum subquadrate, about 1.1 times longer than wide, anterior edge arcuate, sides slightly diverging posteriorly, posterior edge nearly straight, anterior angles rounded, posterior angles nearly rectangular, disc slightly convex at posterolateral parts, surface finely and densely punctate.

Elytra about 3.0 times longer than wide, 4.0 times longer than pronotum, width at humeri greater than posterior edge of pronotum, sides nearly parallel, surface slightly more coarsely and densely punctate than pronotum.

Legs with all tarsal claws bifid, each with lower projection as long as upper one.

Aedeagus (Fig. 2 A–C): ventral process of each paramere wide, slightly narrowed apically and hooked at apex; conjoint dorsal plate of parameres moderately developed, about one-third length of ventral process of each paramere, slightly narrowed apically, with apex medially concave.

Female. Similar to male, but vertex black, antennae black, antenomeres I and II orange, filiform and simple, extending to basal third of elytra. Abdominal sternite VIII (Fig. 4A) nearly straight at posterior edge, lateral angles obtuse.

Internal genitalia (Fig. 5A): vagina abruptly extended apically as a short and thick duct at ventroapical portion; diverticulum and spermathecal duct arising from end of duct of vagina; diverticulum evenly thin, spiraled and long; spermathecal duct slightly shorter and thicker than diverticulum; spermatheca nearly as long as and slightly thicker than diverticulum, with a moderately long thin accessory gland, which is longer than spermatheca.

Body length: 7.9-9.0 mm; width: 1.7-2.0 mm.

Etymology.

The specific name is derived from Latin securiclatus (axe-like), referring to its antennomeres IV–IX securiform in male, which abruptly widened apically and rounded at apical angles.

Diagnosis.

Head mostly orange, pronotum and elytra uniformly black; male antennomeres IV–IX securiform, abruptly widened apically and rounded at apical angles; aedeagus: conjoint dorsal plate of parameres moderately developed, about one-third length of ventral process of each paramere, slightly narrowed apically, with apical edge medially concave.

Remarks.

No other species has the male antennomeres IV–IX securiform as in this species. This species is similar to F. pallidiceps (Pic, 1911) in the body coloration, but can be easily distinguished by the male antennae, of which the antennomeres III–IX are widened apically, while simple filiform in F. pallidiceps ( Wittmer 1988: fig. 6); aedeagus: the conjoint dorsal plate of parameres is slightly narrowed apically, with the apex medially concave, while distinctly narrowed apically, with the apex straight in F. pallidiceps ( Wittmer 1988: fig. 26). It also resembles F. liuchowensis (Wittmer, 1989), but can be differentiated by the male antennae, of which the antennomeres IV–IX are abruptly widened apically and rounded at apical angles, while obliquely widened apically in F. liuchowensis ( Wittmer 1989: fig. 8); aedeagus: the conjoint dorsal plate of parameres is moderately developed, while greatly reduced in F. liuchowensis ( Wittmer 1989: fig. 9).