Tituboea macropus ( Illiger, 1800 )

Bezděk, Jan, 2016, Identity of taxa proposed inClythra (Coleoptera: Chrysomelidae: Cryptocephalinae) by Carl Peter Forsberg (1821), Acta Entomologica Musei Nationalis Pragae 56 (2), pp. 769-784 : 781

Tituboea macropus ( Illiger, 1800)

( Figs 41–44 )

Clytra macropus Illiger, 1800: 128 (original description, senior homonym). Clythra coalita Forsberg, 1821a: 264 , 280 (original description), syn. nov. Clythra grandipes Forsberg, 1821a: 262 (unnecessary new substitute name for Clytra macropus Illiger, 1800 ).

Type localities. Clytra macropus : ‘Friaul’ [= Northeastern Italia / Slovenia, Friuli region]. Clythra coalita : not stated. Clythra grandipes : ‘Germania’.

Type material examined. Clytra macropus : not examined.

Clythra coalita : HOLOTYPE: J, ‘ Uppsala Univ. Zool. Mus. / Thunbergsaml. nr. 8217 / Typsaml. nr. 1758 / Clythra coalita Forsberg 1821 [r, p] // coalita. / α. / 28 [box label, w, h]’ ( UUZM).

Clythra grandipes : not examined.

Distribution. Central and Southeastern Europe, Turkey, Caucasus, Near East, Central Asia (for details see REGALIN & MEDVEDEV 2010).

Comments. LACORDAIRE (1848) placed Clythra coalita among the species not known to him, correctly stating that the type locality was not mentioned in the original description, but mistakenly proposed its origin as from ‘Cap de Bonne Espérance’ [= Cape of Good Hope]. This mistake was repeated in subsequent catalogues ( GEMMINGER & HAROLD 1874, JACOBY & CLAVAREAU 1906, CLAVAREAU 1913) and, evidently, it is the reason why this taxon was not treated by specialists in Palaearctic fauna.

The holotype of Clythra coalita is in very poor condition, the preserved parts being: part of pronotum, elytra, part of thorax, both fore legs and left mid leg. However, there is no doubt about the identity of the holotype and Clythra coalita is proposed as a new junior synonym of Tituboea macropus .

FORSBERG (1821a) proposed the new name Clythra grandipes to replace Clytra macropus Illiger, 1800 . Although this nomenclatural act was not explained, it seems to be evident that Forsberg realized the homonymy of Clytra macropus Illiger, 1800 and Clythra macropus Thunberg, 1821 , but mistakenly replaced the older name of Illiger, instead of the younger name provided by Thunberg. Clythra grandipes should be listed in synonymy with Tituboea macropus , as can be found in all important publications and catalogues ( LACORDAIRE 1848, GEMMINGER & HAROLD 1874, JACOBY & CLAVAREAU 1906, CLAVAREAU 1913, REGALIN & MED- VEDEV 2010).

Figures

Figs 36–44.36–40 – Tituboea biguttata (Olivier, 1791) (holotype of Clythra sexpunctata Forsberg, 1821,♀, 8.5 mm): 36 – dorsal habitus; 37 – lateral habitus; 38 – frontal habitus; 39 – box label; 40 – label of holotype. 41–44 – Tituboea macropus (Illiger, 1800) (holotype of Clythra coalita Forsberg, 1821,J, 10.0 mm): 41 – dorsal habitus; 42 – lateral habitus; 43 – box label; 44 – label of holotype.