Camchaya loloana var. loloana

Bunwong, Sukhonthip, Chantaranothai, Pranom & Keeley, Sterling C., 2014, Revisions and key to the Vernonieae (Compositae) of Thailand, PhytoKeys 37, pp. 25-101 : 52-53

Camchaya loloana var. loloana

Description.

Annual herbs, 30-70 cm tall. Stems erect, terete, inconspicuously ribbed, pubescent with T-shaped hairs and glands. Leaves ovate, 3-10 by 2-4 cm, margin serrate, apex acute or acuminate, base attenuate, chartaceous; both surfaces scabrous with whip-shaped hairs, cylindrical hairs and capitate glands; lateral veins 8-12-paired; petioles up to 10 mm long. Capitulescences terminal and axillary, paniculate. Capitula campanulate, 9-10 mm long, pedunculate. Receptacle convex, 3.5-4 mm in diam., glabrous. Involucres campanulate, in 7-8 series, 8-10 mm long. Phyllaries imbricate, greenish with purple apex, margin with spines up to 1 mm long, outer surface arachnoid, glands capitate; the outer lanceolate, apex spinose; the inner ones lanceolate or linear-oblong, apex acuminate. Florets 65-100; corollas funnelform, purple rarely white, puberulous, glands capitate; corolla tubes 7-9.5 mm long; corolla lobes 2.5-3 mm long. Anthers 1.8-2.5 mm long, apical appendage acute, base rounded. Styles purple, 7-11 mm long, branches ca. 2 mm long. Achenes obovate, 1.3-1.7 mm long, glabrous, 10-ribbed. Pappus in one series, bristles 1.5-3 mm long, absent or deciduous.

Distribution.

Thailand: Chiang Mai, Lampang, Phitsanulok, Khon Kaen, Nakhon Ratchasima, Ubon Ratchathani, Kanchanaburi, Saraburi. China (Yunnan), Laos, Myanmar.

Specimens examined.

Thailand, Khon Kaen, Phu Wiang national park, 16°40.93'N, 102°14.15'E, 1 Oct 2007, S. Bunwong 330 (KKU, US); Chiang Mai, Doi Chiangdao, 9 Nov 1922, A.F.G. Kerr 6650 (BM, K, P); Doi Chiangdao, 30 Jul 1968, K. Larsen, T. Santisuk & E. Warncke 2862 (BKF, E); Doi Chiangdao, 27 Sep 1994, W. Nanakorn et al. 1821 (QBG); Doi Chang, 29 Oct 1979, T. Shimizu, H. Toyokumi, H. Koyama, T. Yahara & T. Santisuk 20639 (AAU, BKF, L); Khon Kaen, 18 Sep 1994, W. Nanakorn et al. 1626 (QBG),9 Jan 1997, W. Nanakorn et al. 8472 (QBG); Ubon Ratchathani, Sirinthon Dam, 27 Oct 2007, S. Bunwong 339 (KKU, US); Kanchanaburi, Kritee, 9 Jul 1973, R. Geesink & C. Phengklai 6192 (AAU, BKF, E, L, P); Sangkhlaburi, 14 Jul 1973, S. Sutheesorn 2637 (BK); Saraburi, Chaibadan, 15 Dec 1923, A.F.G. Kerr 7982 (BK, BM, K, P).

Diagnostic characters.

Camchaya loloana is recognized by having short spines on the margins of the phyllaries, 10-ribbed achenes, and leaves with T-shaped hairs.

Ecology.

On limestone in dipterocarp, dry evergreen, and hill evergreen forest, alt. 400-1500 m; flowering July to December.

Vernacular name.

Dok Lea (ดอกแล่), Phu Muang (พู่ม่วง).