Tentyria

Bujalance, José L., Ferrer, Julio & Cárdenas, Ana M., 2023, A taxonomic revision of the genus Tentyria Latreille, 1802 in the Iberian Peninsula and Balearic Islands (Coleoptera: Tenebrionidae), Zootaxa 5320 (1), pp. 1-88 : 56-61

Key of the species of Tentyria from the Iberian Peninsula, Columbretes islands and Balearic archipelago:

1 Base of pronotum protruding backwards forming a central lobe, bi-dentate or indented, seldom truncate ( Figs.70–77 )..... 2

- Base of pronotum without these features ( Figs. 78–105 )...................................................... 3

2 Pronotum configured according to Figs. 70–72 ; base of elytra entirely margined, excavated only in the middle, near the scutellum, just surrounding the basal lobe of pronotum ( Figs. 143, 144 ); parameres of the aedeagus with the apex truncated or blunt ( Figs. 154–156 )................................................................................ 13

- Pronotum configured according to Figs. 73–77 ; base of elytra backward curved, from the humeri ( Fig. 142 ), which are somewhat angled, but always well noticeable; the margin thin or erased before the scutellum; parameres of the aedeagus with the apex sharp, mucronate or distinctly toothed ( Figs. 157–161 )....................................................... 4

3 Species showing the apex of the anal sternite bifid ( Fig. 153 ).................................................. 25

- Species showing the anal sternite normally configured, rounded or truncate, but not bifid ( Figs. 149–152 )............... 5

4 Head with parallel temples and epistome truncate and usually without perceptible tooth ( Figs. 4–7 ); pronotum finely punctured, teeth of the central lobe very blunt ( Figs. 73–76 ); base of elytra not completely margined; male protibiae commonly sinuous in the inner margin and longer than those of the female ( Fig. 141 ); anal sternite truncated apically ( Fig. 149 ); aedeagus with parameres elongated and of similar length as the phallobase ( Figs. 157–160 )..................................... 14

- Head with the epistome rounded from the genae and provide with a small tooth somewhat perceptible ( Fig. 8 ); pronotum with dense and strong punctures; base fully margined and with the central lobe commonly feebly excavated or bidentate ( Fig. 77 ); base of elytra entirely margined; anal sternite rounded apically ( Fig. 150 ); aedeagus with parameres curved and conspicuously shorter than the phallobase ( Fig. 161 ).................................................................... 15

5 Large head, broad and widely extended in front of eyes which are very convex, epistome triangular, thickened ahead, and with a noticeable tooth in the middle ( Fig. 31 ); gular groove very wide, and transverse and deep ( Fig. 65 ); pronotum with the basal margin curved, very thick and sinuous before the posterior angles ( Fig. 101 ); prosternal apophysis protruding behind the procoxae ( Fig. 136 ); elytra with the basal margin thick and raised, especially in the humeri which are very well marked ( Fig. 145 ); robust and large specimens; aedeagus large and robust, about 5 mm in length ( Fig. 181 )....................... 22

- Without the abovementioned features..................................................................... 6

6 Head with very convex eyes, temples converging to the vertex, epistome somewhat rounded and provided with a small median tooth, more conspicuous in lateral view ( Figs. 15, 16 ); gular groove commonly apparent and more or less deep ( Figs. 50, 51 ); pronotum transverse, almost as wide as the elytra, particularly in males, with dense punctures ( Fig. 84 ); apophysis prosternal commonly protruding backwards, between the procoxae ( Figs. 120, 121 ); inner margin of protibiae in males feebly sinuous, and longer than in females ( Fig.141 ); elytra densely punctured, usually more or less striated; humeri rounded, not angulous; aedeagus shaped as in Figs. 168, 169.................................................................... 18

- Without the abovementioned features together.............................................................. 7

7 Head with the sides subparallel or parallel, temples not converging to vertex, eyes flattened, epistome truncate or sub-truncate, exceptionally rounded, often trapezoidal ( Figs. 9–13 ); gular groove superficial, generally feebly delimited ( Figs. 44–48 ); pronotum with punctures fine and sparse, the base curved and more or less protruding backwards and strongly margined ( Figs. 78–82 ); base of elytra curved, receiving the base of pronotum; apex of the anal urosternite truncated or slightly sinuate ( Fig. 149 ); aedeagus according to Figs. 162–166................................................................. 16

- Head with the epistome more or less rounded anteriorly, triangular or with the margin sinuate ( Figs. 14–35 ); overall, without the abovementioned features together..................................................................... 8

8 Head short, eyes flattened, exceptionally somewhat convex, the supraorbital fold slightly raised; epistome rounded from the genae, with a small tooth at middle ( Figs. 24–27 ); gular groove commonly narrow and normally reduced to a central depression ( Figs. 58–61 ); Pronotum with the base, usually, curved and more or less protruding backwards, finely margined, sometimes slightly sinuate before the posterior angles, which are obtuse and usually not well noticeable, proportionally large, sometimes nearly as broad as the elytra and with dense punctures, the sides regularly curved, but the curve more pronounced at base ( Figs. 92–97 ); protibiae lack clear sexual dimorphism; aedeagus with the parameres shorter or almost equal than the phallobase ( Figs. 176–179 ).......................................................................................... 20

- Without the abovementioned features together.............................................................. 9

9 Head proportionally large, eyes flattened; epistome triangular or subtriangular, and provided with a conspicuous tooth ( Fig. 14 ); gular groove broad, very deep, well delimited at each side ( Fig. 49 ); pronotum very variable with very fine punctures, usually not transverse; base almost straight, never sinuate before posterior angle and often narrower than the anterior margin ( Fig. 83 ); prosternal apophyse elongated, surpassing the procoxae ( Fig. 119 ); elytra narrow, with subparalel sides, slightly broader than pronotum, sometimes with vestigial striae and variable in length; aedeagus with the parameres widened in the middle, slightly longer than phallobase ( Fig. 167 )........................................................................ 17

- Without the abovementioned features together............................................................. 10

10 Head broad, epistome more or less protruding forwards, but never truncate or triangular, with dense punctures, without medial tooth or very inconspicuous, commonly with the maximum width after eyes, because of presenting very convex temples, sometimes become broader before eyes because of the genae are widened, two lateral cephalic depressions at each side, at the level of the antennal insertion, and sometimes with another depression on the disc, supraorbital folds very raised, curved and sinuate, rarely sub-straight ( Figs. 28–30 ); gular groove straight, narrow and moderately deep ( Figs. 62–64 ); pronotum scarcely convex, distinctly transverse, the base broadly margined, curved and often more or less sinuates before posterior angles ( Figs. 98–100 ); elytra smooth or feebly rough, sometimes with vestigial striae; aedeagus with parameres conspicuously longer than phallobase and with slightly sinuate sides; penis narrowed before the apex ( Fig. 180 ).............................. 21

- Without the abovementioned features together............................................................. 11

11 Elytra provided with striae and wrinkles in zigzag; shape oval-elongate from base to apex, which is normally rounded, not bifid, with the base in line almost straight towards the scutellum; Head as broad as long, narrowed behind the eyes, which are large and feebly convex, rounded epistome without tooth in the middle, the supra-orbital fold conspicuous ( Fig. 34 ); pronotum convex and feebly transverse, base broadly margined, protruding in curved backwards, feebly sinuate before the posterior angles, which are obtuse, the punctures strong and dense, similar to those of the head ( Fig. 104 ); aedeagus presenting the parameres slightly longer than the phallobase and slightly sinuate ( Fig. 184 ); matte tegument................................. 24

- Without the abovementioned features together............................................................. 12

12 Pronotum transverse, slightly convex with the sides openly curved from base to the apex, base curved and barely protruding backwards, normally slightly sinuate before posterior angles, which are obtuse and not or barely indicated ( Figs. 102, 103 ); gular groove narrow, shallow and poorly defined in the centre ( Figs. 66, 67 ); male protibiae quite similar to those of female; aedeagus narrow and elongate, the parameres slightly longer or clearly longer than the phallobase ( Figs. 182, 183 ); tegument smooth and shiny.................................................................................... 23

- Pronotum moderately or slightly transverse, in the latter case very convex, with the base straight or slightly curved and somewhat protruding backward ( Figs. 85–91 ); usually without the abovementioned features together.......................... 19

13 GROUP OF Tentyria mucronata (Northeastern Iberian Peninsula and Balear archipelago):

A Basal lobe of pronotum with the margin thinned or deleted in the middle, often bimucronate ( Fig. 70 ); humeri rounded, the base of the elytra narrowed and very curved from the humerus ( Fig. 144 ); aedeagus narrow, about 3.7 mm in length, with the parameres somewhat longer than the phallobase, the side subparallel and sinuate ( Fig. 154 ); species from the northeastern coast of the Iberian Peninsula...................................................................... T. mucronata

- Basal lobe of pronotum with the margin more or less thick, slightly bimucronate or truncate ( Figs. 71, 72 ); base of the elytra wide, curved next to the scutellum and in straight line till the humeri which are angled and fallen down ( Fig. 143 ); aedeagus proportionally wider, and with sides less parallel than in the previous species ( Figs. 155, 156 ); species from the Balearic archipielago......................................................................................... B

B Head with the eyes slightly convex, not clearly exceeding the lateral contour, epistome broad and rounded at its leading edge ( Fig. 2 ); pronotum transverse, the sides regularly curved from the basis to the apex, with the basal lobe scarcely back protruding, very slightly bidentate ( Fig. 71 ); Elytra striated or rough-striated and broader than the pronotum; body robust and tegument dull................................................................................ T. schaumi

- Head with big and convex eyes, exceeding the lateral contour, epistome narrower and subtriangular ( Fig. 3 ); pronotum less transverse, with narrower sides towards the base than towards the apex; base of pronotum narrower and margined, with the middle lobe barely indented, rather truncated or even rounded ( Fig. 72 ); elytra barely broader than pronotum, smooth or weakly striate and very oblong in appearance; graceful and bright body, with very fine and hardly noticeable punctures.. T. ophiusae

14 GROUP of Tentyria subcostata (littoral the meridional Iberian Peninsula):

A Base of pronotum entirely margined, sinuate before the posterior angles, which are straight or sub-straight and well-marked ( Figs. 74, 75 ).............................................................................. T. sinuatocollis

a Pronotum barely transverse, at least in males, maximum width at the middle, punctures fine and sparse and the basis noticeaably trisinuated ( Fig. 74 ); elytra ovate and elongate, widener in females....................... ssp. sinuatocollis

- Pronotum transverse, subcordiform, maximum width before the middle, the punctures almost imperceptible, the base usually bisinuous and the medium lobe frequently truncate ( Fig. 75 ); elytra shorter and broader, often somewhat deep at the suture line.................................................................................. ssp. escalerai nov.

- Base of pronotum with the margin very fine and, commonly deleted at the medium lobe, posterior angles very obtuse, barely indicated in dorsal view ( Figs. 73, 76 )................................................................... B

B Pronotum transverse and convex, maximum width at the middle, constricted forwards and backwards, resulting in a nearly pentagonal shape ( Fig. 76 ); sexual dimorphism of protibiae not observable; elytra broad and convex, variably striated or with vestigial striae; aedeagus with parameres as long and broad as the phallobase, endophallus (penis) thick but sharply constricted before apex ( Fig. 160 ).......................................................................... T. lateritia

- Pronotum moderately convex, the sides regularly curved from the base to the apex ( Fig. 73 ); protibiae in males long and sinuate in the inner side; overall, without the abovementioned features................................................ C

C Elytra robust and conspicuously ribbed and rough, with fine or inconspicuous punctures; the punctures in the pronotum fine but well distinguishable; aedeagus of 3.5-4 mm length, with the parameres strongly acuminate at the apex ( Fig. 158 ); usually large and robust specimens...................................................................... T. striatorugosa

- Elytra smooth, or with vestigial ribs, finely but more conspicuously punctured than the previous species; the punctures in the pronotum less noticeable; aedeagus smaller, about 3–3.5 mm length, and with the parameres not so strongly acuminate ( Fig. 157 ); smaller and more graceful specimens...................................................... T. subcostata

15 GROUP OF Tentyria faroensis (Southern Portugal):......................................... T. faroensis sp. nov.

16 GROUP OF Tentyria curculionoides (Atlantic, Western and Northwestern species):

A Supraorbital folds generally feebly elevate and separate from the eye in dorsal view ( Figs. 9, 10 ); pronotum with the posterior angles obtuse and usually inconspicuous in dorsal view, the base backwards prolonged continuing the curvature of the sides of the pronotum ( Figs. 78, 79 ); elytra generally very convex and robust, rough-striated very finely punctured, provide with wide and slightly elevated ribs, crossed by transverse wrinkles; base of the elytra fully margined; sexual dimorphism of protibia unremarkable; aedeagus with regularly narrowed parameres from the base to the apex, of similar or shorter length than phallobase, the endophallus broad as in T. lateritia ( Figs. 162, 163 )............................................. B

- Supraorbital fold well developed and next to the eyes, the space between them invisible in dorsal view ( Figs. 11–13 ); pronotum with posterior angles obtuse and generally well indicated, the base openly curved, scarcely protruding backwards and often sinuated at the posterior angles ( Figs. 80–82 ); elytra smooth or only with vestigial lines of striae, and punctures as the pronotum, frequently with the basal margin interrupted before the scutellum; sexual dimorphism of protibia is conspicuous; aedeagus more stylized with the parameres subparallels and similar in length or longer than the phallobase ( Figs. 164–166 )....... C

B Pronotum transverse, subpentagonal, basal margin broad and slightly sinuate to the posterior angles ( Fig. 79 ); prosternal apophysis narrow and lanceolate ( Fig. 115 ); aedeagus with parameres nearly equal or shorter than the phallobase ( Fig. 163 )................................................................................... T. stupefacta sp. nov.

- Pronotum more regularly curved than in the precedent species, not subpentagonal, basal margin with finer carena, forming a regular curve, not so protruding backwards ( Fig. 78 ); prosternal apophyses broadened and rounded at the end, not lancelolate ( Fig. 114 ); aedeagus more robust, with the parameres and the phallobase very similar in length ( Fig. 162 ). T. curculionoides

C Supraorbital fold noticeable, with straight margin ( Fig. 11 ); pronotum with the sides in a more closed curve towards the base than at the apex, the sides and the base slightly sinuated before the posterior angles which are well indicated, base curved and protruding backwards ( Fig. 80 ); elytra generally smooth or with vestigial striae, base of elytra entirely margined. aedeagus reaching about 3.8 mm length, with the parameres slightly longer than the phallobase ( Fig. 164 ) ............... T. heydeni

- Supraorbital fold barely noticeable, with sinuous margin ( Figs. 12, 13 ); pronotum regularly and openly curved at sides, from base to anterior angles, not sinuate before posterior angles, base not so protruding backwards ( Figs. 81, 82 ); base of elytra with the margin interrupted or rough and fuzzy before the scutellum; elytra more oval-elongate, not so convex and barely wider than pronotum, often striated-rough.......................................................................... D

D Head with epistome truncate ( Fig. 12 ); pronotum convex, broad and transverse, with well indicated posterior angles ( Fig. 81 ); males protibiae almost equal to those of the female; elytra frequently very rough-striated, punctation greater than those of the pronotum, with the humeri well indicated; aedeagus reaching about 4.3 mm lenght, the parameres conspicuously longer (1/3 times) than the phallobase ( Fig. 165 ); usually large and robust specimens with shiny black tegument.... T. espanoli sp. nov.

- Head with epistome in general sub-truncate and longer ( Fig. 13 ); pronotum barely convex, with posterior angles not so conspicuous ( Fig. 82 ); males with longer and slender protibiae; elytra slightly constricted in the base, superficially striate-rough and sometimes practically smooth, very fine punctures, similar or less than those of the pronotum, the humeri almost rounded, non-angled; aedeagus of smaller size and with the parameres nearly equal or slightly longer than the phallobase ( Fig. 166 ); specimens more graceful and smaller, with matte tegument and black, but less intense..................... T. interrupta

17 G ROUP OF Tentyria platyceps (species from the interior centro-meridional lands of Iberian Peninsula):....... T. platyceps

18 GROUP OF Tentyria bassii (species from the interior of the Iberian Peninsula, distributed in the western half, especially in the southwestern quadrant): A Apex of elytra very acuminated or spiniform due to the convergence of the upper and lower margins of the epipleura ( Fig. 146 ); broad and convex eyes ( Fig. 16 )............................................................... T. eulipoides

- Apex of elytra not bifid neither spiny; the head is narrower than in the previous species, with smaller eyes ( Fig. 15 ); females usually with very stubby elytra; tegument with more strong and denser punctures; smaller and more graceful specimens ….…............................................................................................. T. bassii

19 GROUP OF Tentyria peiroleri (south-eastern Iberian Peninsula):

A Head showing large eyes, with sharp and denticulate epistome ( Fig. 22 ); gular groove wide and deep, well defined ( Fig. 56 ), comparable to T. grossa ssp. basalis and T. platyceps ; pronotum barely transverse, at least in males, very convex, sub-globous, base straight or slightly curved ( Fig. 90 ), strong and dense, but not confluent punctures; prosternal apophysis with the end rounded and usually broadly margined ( Fig. 127 ); parameres with the sides slightly narrowing from the base to the apex, and barely longer than the phallobase which is oval shaped ( Fig. 174 )................................... T. kochi sp. nov.

- Gular groove narrowed, at least on the sides, variable in depth, but never like in the previous species ( Figs. 52–55 ); in general, without the previous features combined.................................................................. B

B Head with the epistome of sub-rounded to angled, but always with a more or less developed tooth ( Figs. 17, 18 ); gular groove moderately deep and often openly “V” shaped ( Fig. 52 ); pronotum convex, scarcely transverse or sub-globose ( Figs. 85, 86 ), with fine but dense and well-marked punctures almost equal to that of the head; short and oval elytra, on average less than 2.75 times the length of the prothorax; often more chubby in females; sometimes with signs of stretch marks and finely punctuated; apex non spiniform; aedeagous of 2.9–3.6 mm long, with the parameres subparallel at the sides, slightly narrower than the phallobase, which is similar in length or somewhat smaller than the parameres ( Fig. 170 ); specimens, usually, not exceeding 14 mm in length................................................................................. T. peiroleri

a Head broad, epistome barely angulate or subrounded and slightly sinuate at the sides ( Fig. 17 ); pronotum very slightly transverse, the sides regularly curved, somewhat closed towards the base, obtuse and very slightly indicated posterior angles, the basal margin almost straight and barely sinuate before the posterior angles ( Fig. 85 ); prosternal apophyse broad and barely prolonged beyond the procoxas ( Fig. 122 ); oval elytra, often very chubby in females; last abdominal sternite slightly sinuate at the apex ( Fig. 151 ). Specimens with an average size of 14 mm in length............................... ssp. peiroleri

- Narrower head, usually with angulate or subtriangular epistome, ( Fig. 18 ); pronotum less transverse and more convex, frequently suborbicular ( Fig. 86 ); prosternal apophyses narrower and elongated, often pointed and surpassing the level of the procoxas ( Fig. 123 ); last abdominal sternite not or barely sinuate at the apex; elytra proportionally shorter and narrower, often sub-cylindrical; smaller specimens, average 13.3 mm............................................... ssp. incerta

- Without the above characteristics combined............................................................... C

C Head with the epistome somewhat projected forward, rounded or sub-truncated without perceptible tooth ( Figs. 19, 23 ); the gular groove ( Figs. 53, 57 ) always very well noticeable and occasionally broad and deep, although never comparable to that of T. kochi ; pronotum slightly transverse in males, well punctured, with the sides in open curve and anterior angles well indicated, the base is straight or slightly curved and generally sinuated at the posterior angles ( Figs. 87 , 91 ); more transverse and convex in females; elytra almost equally narrowed at the basis than at the apex; aedeagus with the parameres usually exceeding 2 mm length, and always longer and narrower than the phallobase ( Figs. 171 , 175 )..................................... D

- Head with rounder epistome and, often provided of a tiny tooth at the apex ( Figs. 20, 21 ); the gular groove consisting in a central depression or narrow and shallow line ( Figs. 54, 55 ). Pronotum more transverse, with sides in closer curve and the anterior angles rounded ( Figs. 88, 89 ), occasionally sub-circular in shape. Wide and oval elytra and clearly narrower towards the apex, which can be spiny; parameres with the maximum width in the middle, almost as broad as the phallobase and a little longer than this, usually not more than 2 mm in length ( Figs. 172, 173 )................................. T. sublaevis a Head small ( Fig. 20 ); gular groove often consisting of a shallow central depression and two lateral notches ( Fig. 54 ); pronotum transverse, usually with fine punctures ( Fig. 88 ); oval elytra elongated towards the apex, that never is spiny, but often of subparallel sides; aedeagus about 3.4 mm long on average ( Fig. 172 ); specimens graceful, about 13.7 mm long....... ssp. sublaevis

- Big head ( Fig. 21 ); narrow and scarcely deep gular groove ( Fig. 55 ); pronotum variable and somewhat less transverse ( Fig. 89 ); occasionally subcircular; punctures fine but evident and scarcely marked on the disc; oval and elongate elytra, and clearly narrower towards the apex, which is acuminate or even spiny by the prolongation of the upper margin of the epipleura, usually exceeding the last abdominal sternite ( Fig. 147 ); aedeagus about 4 mm long on average ( Fig. 173 ). Body broader and more robust, reaching 16 mm long............................................................... ssp. cognata nov.

D Pronotum densely punctured, small, about 2.8–3.3 mm long and 3.5–4.5 mm width, with the basis curved and usually sinuate before the posterior angles ( Fig. 87 ); oval and elongated elytra, nearly three times longer than pronotum; the parameres of 1.1 times longer, on average, than the phallobase ( Fig. 171 ); graceful specimens, 14.5 mm medium lengt.......... .. T. prolixa

- Pronotum of greater size (3.3–4 mm long; 4.3–5.2 mm broad) and less densely punctured ( Fig. 91 ); elytra oval, shorter and wider, on average 2.75 times longer than the pronotum, with the apex somewhat sharp but not spiniform; aedeagus of greater size, about 4.25 mm long, with the parameres 1.25 times longer, on average, than the phallobase ( Fig. 175 ); specimens more robust and shinier,15.4 mm medium length............................................... T. castrotovari sp. nov.

20 GROUP OF Tentyria gaditana (littoral or sub-littoral areas of the southern Iberian Peninsula: Málaga, Cádiz and Huelva):

A Striated-rough elytra, intervals slightly elevated and with tuberous wrinkles; pronotum transverse, with maximum width before the middle, very curved sides and narrowing towards the base which is very extended backwards in some specimens ( Figs. 93, 94 ); protibiae of males slightly sinuate on inner edge; aedeagus robust, with the phallobase equal or nearly as long as the parameres ( Fig. 177 )........................................................................ T. corrugata

- Smooth or sub-striated elytra, but never ribbed; pronotum generally less transverse, with less curved sides and narrowed almost equally towards both ends ( Figs. 92, 95–97 ); protibiae without perceptible sexual dimorfism; aedeagus generally with parameres shorter than phallobase ( Figs. 176, 178–179 )....................................................... B

B Elytra with narrow and curved base, tapering posteriorly by prolongation of the upper margin of the epipleure beyond the last abdominal sternite, smooth or with signs of striae, frequently depressed in the suture and finely punctured; pronotum fine but densely punctured; aedeagus with the parameres clearly shorter than the phallobase and convergent towards the apex in a straight or sinuous line ( Figs. 178, 179 )................................................................... C

- Elytra smooth, regularly oval from base to apex, not narrowed in the humeri nor acute at the apex, frequently shorter in relation to pronotum, more conspicuously punctured, the punctures sometimes as strong as in the pronotum; pronotum always matt, strong and densely punctured; aedeagus with more parallel sides, with the phallobase similar in length to the parameres ( Fig. 176........................................................................................ T . gaditana

C Pronotum not very convex, with the base protruding backwards, narrower than the apex, and often sinuous before of the posterior angles ( Figs. 95, 96 ); elytra oval elongated often sub-striate; parameres of aedeagus with the sides not sinuous ( Fig. 178 )........................................................................................................................................................................................ T. donanensis

- Pronotum more transverse and convex, matte; punctures thicker and denser, almost like in the head; base of the pronotum as wide as in the apex, not or barely sinuate front of the posterior angles, barely extended back ( Fig. 97 ); elytra smooth, often stockier, and less constricted in humeri; aedeagus somewhat longer than in the previous species, with the parameters constricted at the base and converging in a straight or slightly sinuous line towards the apex ( Fig. 179 )...... T. pseudogaditana sp. nov.

21 GROUP OF Tentyria velox (provinces of Castilla and Leon):............................................ T. velox

a Elytra with tuberous wrinkles, with aspect of cerebral circumvolutions; pronotum transverse on males, finely punctured, with the base clearly sinuated before the posterior angles and curving backwards, at least in males ( Fig. 99 )....................................................................................... s sp. circumvoluta nom. and stat. nov.

- Elytra smooth or only with vestigial striae or rugosities, never with tuberous wrinkles............................ b

b Pronotum transverse, particularly on males, finely and sparsely punctured, base slightly protruding backwards ( Fig. 98 ); elytra short, oval, finely punctured as pronotum or smaller; specimens in general of small size and tegument smooth and shiny........................................................................................ ssp. velox

- Pronotum less transverse, at least on males, with dense and strong, but not confluent punctures; the base more curved backwards than in the previous species ( Fig. 100 ); elytra long and oval shaped, often slightly prolongate beyond the last abdominal sternite, densely punctured, similar in size as the pronotum and the head. Body broad and robust; tegument less shiny or matte........................................................................... s sp. serrana nov.

22 GROUP OF Tentyria grossa (Balearic Islands):............................................. T. grossa ssp. basalis

23 GROUP OF Tentyria castiliana (Center-Oriental regions of the Iberian Peninsula):

A Head with eyes barely convex, epistome rounded anteriorly and slightly sinuate at sides ( Fig. 32 ); pronotum generally quite transverse, sides in narrower curve toward the base ( Fig. 102 ); apex of each elytra thorn shaped, sometimes not very observable, by fusion of the inferior margin of the epipleura that ends fused to the superior margin ( Fig. 148 ); sexual dimorphism in protibiae almost unperceivable; aedeagus stylized, long and subparallel, with similar or barely longer parameres than the phallobase ( Fig. 182 )......................................................................... T. castiliana

- Head with more convex eyes and epistome extended ahead, well separated from the genae by a conspicuous sinuosity ( Fig. 33 ); pronotum less transverse, stronger, and densely punctured, with the base more curved and sinuate before the posterior angles ( Fig. 103 ); apex of elytra not spiniform; protibiae in male feebly sinuous in the inner margin and longer than in females; aedeagus with the parameres clearly longer than the phallobase ( Fig. 183 ).............................. T. aragonica

24 GROUP OF Tentyria pazi (Columbretes Islands) :....................................................... T. pazi

25 GROUP OF Tentyria bifida (Southwestern distribution, Doñana National Park):............................ T. bifida

Figures

FIGURES 70–87. I / Pronota I. 70: T. mucronata. 71: T. schaumii. 72: T. ophiusae. 73: T. subcostata / T. striatorugosa. 74: T. sinuatocollis ssp. sinuatocollis. 75: T. sinuatocollis ssp. escalerai. 76: T. lateritia. 77: T. faroensis. 78: T. curculionoides. 79: T. stupefacta. 80: T. heydeni. 81: T. espanoli. 82: T. interrupta. 83: T. platyceps. 84: T. bassii / T. eulipoides. 85: T. peiroleri ssp. peiroleri. 86: T. peiroleri ssp. incerta. 87: T. prolixa.

FIGURES 88–105. II / Pronota II. 88: T. sublaevis ssp. sublaevis. 89: T. sublaevis ssp. cognata. 90: T. kochi. 91: T. castrotovari. 92: T. gaditana. 93, 94: T. corrugata. 95, 96: T. donanensis. 97: T. pseudogaditana. 98: T. velox ssp. velox. 99: T. velox ssp. circumvoluta. 100: T. velox ssp. serrana. 101: T. grossa ssp. basalis. 102: T. castiliana. 103: T. aragonica. 104: T. pazi. 105: T. bifida.

FIGURES 141–153. Figure 141: Protibiae of T. eulipoides (a: ♁, b: ♀). Figures 142–145: Base of the elytra. 142: T. striatorugosa / T. sinuatocollis. 143: T. ophiusae. 144: T. mucronata. 145: T. grossa ssp. basalis.

FIGURES 154–171. I. / Aedeagi I. 154: T. mucronata. 155: T. schaumii. 156: T. ophiusae. 157: T. subcostata. 158: T. striatorugosa. 159: T. sinuatocollis f.t. 160: T. lateritia. 161: T. faroensis. 162: T. curculionoides. 163: T. stupefacta. 164: T. heydeni. 165: T. espanoli. 166: T. interrupta. 167: T. platyceps. 168: T. bassii. 169: T. eulipoides. 170: T. peiroleri. 171: T. prolixa.

FIGURES 1–35. Heads. 1: T. mucronata. 2: T. schaumii. 3: T. ophiusae. 4: T. subcostata. 5: T. striatorugosa. 6: T. sinuatocollis. 7: T. lateritia. 8: T. faroensis. 9: T. curculionoides. 10: T. stupefacta. 11: T. heydeni. 12: T. espanoli. 13: T. interrupta. 14: T. platyceps. 15: T. bassii. 16: T. eulipoides. 17: T. peiroleri ssp. peiroleri. 18: T. peiroleri ssp. incerta. 19: T. prolixa. 20: T. sublaevis ssp. sublaevis. 21: T. sublaevis ssp. cognata. 22: T. kochi 23: T. castrotovari. 24: T. gaditana. 25: T. corrugata. 26: T. donanensis. 27: T. pseudogaditana. 28: T. velox ssp. velox. 29: T. velox ssp. circumvoluta. 30: T. velox ssp. serrana. 31: T. grossa ssp. basalis. 32: T. castiliana. 33: T. aragonica. 34: T. pazi. 35: T. bifida.

FIGURES 36–69. Gular grooves. 36: T. mucronata. 37: T. schaumii. 38: T. ophiusae. 39: T. subcostata. 40: T. striatorugosa. 41: T. sinuatocollis. 42: T. lateritia. 43: T. faroensis. 44: T. curculionoides. 45: T. stupefacta. 46: T. heydeni. 47: T. espanoli. 48: T. interrupta. 49: T. platyceps. 50: T. bassii. 51: T. eulipoides. 52: T. peiroleri. 53: T. prolixa. 54: T. sublaevis ssp. sublaevis. 55: T. sublaevis ssp. cognata. 56: T. kochi. 57: T. castrotovari. 58: T. gaditana. 59: T. corrugata. 60: T. donanensis. 61: T. pseudogaditana. 62: T. velox ssp. velox. 63: T. velox ssp. circumvoluta. 64: T. velox ssp. serrana. 65: T. grossa ssp. basalis. 66: T. castiliana. 67: T. aragonica. 68: T. pazi. 69: T. bifida.

FIGURES 106–140. Prosternal apophyses. 106: T. mucronata. 107: T. schaumii. 108: T. ophiusae. 109: T. subcostata. 110: T. striatorugosa. 111: T. sinuatocollis. 112: T. lateritia. 113: T. faroensis. 114: T. curculionoides. 115: T. stupefacta. 116: T. heydeni. 117: T. espanoli. 118: T. interrupta. 119: T. platyceps. 120: T. bassii. 121: T. eulipoides. 122: T. peiroleri ssp. peiroleri. 123: T. peiroleri ssp. incerta. 124: T. prolixa. 125: T. sublaevis ssp. sublaevis. 126: T. sublaevis ssp. cognata. 127: T. kochi. 128: T. castrotovari. 129: T. gaditana. 130: T. corrugata. 131: T. donanensis. 132: T. pseudogaditana. 133: T. velox ssp. velox. 134: T. velox ssp. circumvoluta. 135: T. velox ssp. serrana. 136: T. grossa ssp. basalis. 137: T. castiliana. 138: T. aragonica. 139: T. pazi. 140: T. bifida (a ventral view; b. lateral view).

FIGURES 172–185. II. / Aedeagi II. 172: T. sublaevis ssp. sublaevis. 173: T. sublaevis ssp. cognata. 174: T. kochi. 175: T. castrotovari. 176: T. gaditana. 177: T. corrugata. 178: T. donanensis. 179: T. pseudogaditana. 180: T. velox. 181: T. grossa ssp. basalis. 182: T. castiliana. 183: T. aragonica. 184: T. pazi. 185: T. bifida.