Sagriva Spinola, 1850

Rédei, Dávid, 2017, A revision of Sagriva (Hemiptera: Heteroptera: Dinidoridae), Acta Entomologica Musei Nationalis Pragae 57 (1), pp. 73-95 : 74-78

Sagriva Spinola, 1850

Sagriva Spinola, 1850a: 33, 1850b: 75 . Type species by original designation: Sagriva vittata Spinola, 1850 .

Atelides Dallas, 1852a: 360 . Type species by original designation: Atelides centrolineatus Dallas, 1852 (= Sagriva vittata Spinola, 1850 ). Synonymized by DISTANT (1902: 288) (doubtfully) and BERGROTH (1904: 37).

Sugriva: CASALE (1981: 49). Incorrect subsequent spelling.

Sagriva: SPINOLA (1852a) : 73 (in key), SPINOLA (1852b): 115 (re-publication of original description), STÅL (1876): 127 (listed), LETHIERRY & SEVERIN (1893): 240 (catalogue), BERGROTH (1904): 37 (synonymy), BERGROTH (1908): 188 (listed), KIRKALDY (1909): 260 (catalogue), KIRKALDY (1913): 84 (in key), SCHOUTEDEN (1913): 3, 10 (in key, redescription, distribution, catalogue), HOFFMANN (1948): 25 (catalogue), DURAI (1987): 170, 236 (in key, redescription, revision), LIS (1990): 142 (listed), ROLSTON et al. (1996): 61 (catalogue, bibliography), KOCOREK & DANIELCZOK- DEMSKA (2002): 93 (spermatheca), SCHUH & SLATER (1995): 226 (microptery); GRAZIA et al. (2008): 12 (antenna), LIS et al. (2015): 618 (systematic placement).

Atelides: STÅL (1868) : 522 (in key), WALKER (1868): 500 (listed), STÅL (1870): 89 (listed), STÅL (1876): 127 (listed), ATKINSON (1889): 94 (redescription, fauna of India), LETHIERRY & SEVERIN (1893): 240 (catalogue), DISTANT (1902): 279, 288 (in key, synonymy, redescription, fauna of British India), AHMAD & KHAN (1979): 4, 8 (in key, listed), DATTA et al. (1985): 5 (listed).

Diagnosis. Sagriva is recognized based on the combination of the following characters: mandibular plates far surpassing apex of clypeus, contiguous anteriad of it; lateral margin of head with an angular, transversely directed projection anteriad to eye; eye subpedunculate; antenna 4-segmented, pedicel and basiflagellum angular in cross-section, all faces deeply furrowed between the edges; pronotum subquadrate, anterior margin much broader than head; brachyptery frequent; lateral margins and posterolateral angles of pregenital abdominal segments without tubercles. Detailed redescriptions of the genus were given by SCHOUTEDEN (1913) and DURAI (1987).

Remarks. The title of the paper of DALLAS (1852a), “Description of a new hemipterous insect forming the type [emphasis added] of a new genus”, is considered as an explicit designation of A. centrolineatus as the type species of Atelides under Article 68.2 of the International Code of Zoological Nomenclature ( ICZN 1999).

DURAI (1987) included in the diagnosis of Sagriva that the spiracles of abdominal segment II are concealed by the “metasternum” (correctly metapleuron). This character was found variable during the present study, individuals of both species with completely concealed, partly exposed, and fully exposed spiracles were seen (cf. Figs 17 and 21 ). This phenomenon is apparently due to the fact that the thoraco-abdominal junction allows certain movements between these two tagmata. This character therefore should be used with reservations.

Distribution and diversity. The genus is distributed from Indo-China and the neighbouring areas of the Eastern Himalayas to the western part of the Malay Archipelago. Two species are recognized.

Figures

Figs 13–21. Diagnostic characters of Sagriva vittata Spinola, 1850 (Figs 13–17) and S. banna sp. nov. (Figs 18–21) (brachypterous females). 13, 14, 18 – head, dorsal view; 15, 19 – antenna; 16, 20 – pronotum, scutellum and fore wings, dorsal view; 17, 21 – pterothoracic pleuron, hypocostal lamina, and abdominal ventrite II, most exposed view, evaporatorium densely dotted. Scale bars in mm. Lettering: hcl = hypocostal lamina; sp2 = spiracle of abdominal segment II; v2 = abdominal ventrite II.