Toccolus kuryi, Zhang & Martens, 2020

Zhang, Chao & Martens, Jochen, 2020, A taxonomic study on Epedanidae from Thailand including functional aspects of male genital morphology (Opiliones, Laniatores), ZooKeys 915, pp. 25-58 : 25

publication ID	https://dx.doi.org/10.3897/zookeys.915.47626
publication LSID	lsid:zoobank.org:pub:EA75DB91-3C37-4037-A242-C6F1C144F7CC
persistent identifier	https://treatment.plazi.org/id/F49E92F7-8667-435F-A842-438280CEE8CA
taxon LSID	lsid:zoobank.org:act:F49E92F7-8667-435F-A842-438280CEE8CA
treatment provided by	ZooKeys by Pensoft
scientific name	Toccolus kuryi
status	sp. nov.

Treatment

Toccolus kuryi sp. nov. Figs 102-109 View Figures 102–109 , 110-117 View Figures 110–117 , 118-123 View Figures 118–123 , 124-129 View Figures 124–129

Type material.

Holotype male (SMF-CJM7065): THAILAND: Loel Province: Nang Hin District, near cave Wat Tham Malohan (No.18a), 17°06.47'N 101°52.69'E, alt. 362 m, 2 November 2010, S. Dashdamirov leg. Paratype. One female (SMF-CJM7063): THAILAND: Nong Khai Province: Sangkhom District, Tham Tip Waterfall (No. 23), 18°07.57'N 102°11.16'E, alt. 198 m, 3 November 2010, S. Dashdamirov leg.

Diagnosis.

The distal part of glans nearly triangular (ventral and dorsal views, Figs 119 View Figures 118–123 , 121 View Figures 118–123 ) and the tip curved ventrally (lateral view, Fig. 120 View Figures 118–123 ). Basichelicerite dorsally with three spines (Figs 103 View Figures 102–109 , 104 View Figures 102–109 , 112 View Figures 110–117 , 113 View Figures 110–117 ). Pedipalpal femur ventrally with a row of six setiferous tubercles, the longest one at the base (Figs 108 View Figures 102–109 , 116 View Figures 110–117 ).

Etymology.

The specific name is a patronym in honor of Dr. Adriano B. Kury, a well-known arachnologist from Brazil. A. Kury (2000) created a website of Opiliones and built a platform for acquiring knowledge accessible for everyone. He redefined the family Epedanidae to include four subfamilies ( Kury 1993).

Description.

Male (holotype) habitus as in Figs 102 View Figures 102–109 , 110 View Figures 110–117 , 124-126 View Figures 124–129 . Coloration (Figs 124-126 View Figures 124–129 ): body light yellow with brown patches; median area of prosoma with brown reticulations; both lateral ridges of scutum with brown stripes; opisthosomal areae I-III each with a transverse brownish band; area IV with a series of three brown flecks; all free tergites brown; all coxae yellow with brown reticulations distally; free sternites each with brown bands, these somewhat paler on the central portion; chelicerae and pedipalps reticulated; trochanters pale yellow, femur, patella, tibia and metatarsus with black reticulations, tarsus lighter.

Dorsum (Figs 110 View Figures 110–117 , 124 View Figures 124–129 ). Scutum trapezoid in appearance, both parts nearly parallel-sided, the widest portion of body at opisthosomal area IV, posterior end of opisthosoma bluntly rounded. Prosoma with a row of ten sharply pointed tubercles along anterior margin. Dorsal scutum smooth. Ocularium oval, removed from anterior border of prosoma scutum by 0.38 mm, armed with a conspicuous short median spine of about the height of ocularium (lateral view), four small tubercles above both eyes. The area behind ocularium raised, and the posterior margin of prosoma smoothly rounded. The anterior margin of opisthosomal area I markedly bowed out, borders of other scutal areae slightly bowed out. Opisthosomal areae I-IV with a row of seta-tipped tubercles each and a longitudinal row of similar tubercles on left and right lateral margins. Free tergites with a transverse row of seta-tipped tubercles and anal operculum with scattered granules.

Venter (Fig. 126 View Figures 124–129 ). Coxae with tubercular surface, coxa I additionally with enlarged seta-tipped tubercles. Coxae II-IV with minute tubercles. Genital operculum and free sternites with seta-tipped granules. Spiracles clearly visible.

Chelicerae (Figs 103-106 View Figures 102–109 ). Basichelicerite elongate, armed with two long, curved spines, one short spine dorsally; another one distally on the ectal side; six seta-tipped tubercles scattered over the ventral side; medial side with a basal protuberance (Fig. 103 View Figures 102–109 ). Cheliceral hand considerably widened, pro-dorsally with few seta-tipped tubercles, the inner ones larger than the outer ones; ectally with two seta-tipped tubercles at the subdistal portion; one enlarged seta-tipped tubercle and a smaller one on the ventral side. Fingers strong, inner edges toothed (Fig. 106 View Figures 102–109 ): moveable finger with five teeth, the proximal one cut rectangularly, the other four pointed; fixed finger with five pointed teeth.

Pedipalps (Figs 107-109 View Figures 102–109 ). Coxa dorsally with three seta-tipped tubercles, one strong distal, one sub-distal exterior to it, and a blunt one situated proximally; ventrally with two seta-tipped tubercles. Trochanter ventrally with two setiferous tubercles and dorsally with an enlarge one and a small one. Femur ventrally with a row of five (right pedipalp, Fig. 102 View Figures 102–109 ) or six (left pedipalp, Fig. 108 View Figures 102–109 ) setiferous tubercles, the longest one at the base of the row; dorsally with a row of ten setiferous tubercles, distal one and proximal one inconspicuous; disto-medially side with two setiferous tubercles. Patella ventro-mesally with two and ventro-ectally with one setiferous tubercles. Tibia ventro-mesally with three and ventro-ectally with four setiferous tubercles. Tarsus with three setiferous tubercles each ventro-ectally and ventro-medially. Tarsal claw massive, curved, longer than tarsus.

Legs. All segments unarmed, nearly smooth. Legs I-II slender, legs III-IV strong. Femora I-III not curved, almost straight, femur IV slightly curved. Distitarsi I and II with two tarsomeres. Distitarsi III and IV without scopula and with two unpectinated claws. Tarsal formula (I-IV): 7(2)/16(2)/7/7.

Penis (Figs 118-121 View Figures 118–123 ). Shaft slender, nearly parallel-sided, apical part (pars distalis) distended. Ventral plate conspicuously constricted apically, and dorsally with deep indented near the glans. Glans partially sunken into dorsal depressed portion of pars distalis and extended the distal margin of the ventral plate (Fig. 120 View Figures 118–123 ). The distal part of glans almost triangular (ventral and dorsal views, Figs 119 View Figures 118–123 , 121 View Figures 118–123 ), the tip curved ventrally. Spination symmetrical. One pair of setae A, B, and F. Two pairs of setae C1-2, D1-2, and E1-2 (Figs 120 View Figures 118–123 , 121 View Figures 118–123 ).

Female

(SMF-CJM7063) (Figs 111-117 View Figures 110–117 , 122 View Figures 118–123 , 123 View Figures 118–123 , 127-129 View Figures 124–129 ). Body nearly oval. Granulation and spination of body similar to the male (Figs 111 View Figures 110–117 , 127 View Figures 124–129 ). Prosoma with a row of eight sharp pointed tubercles along anterior margin (Fig. 111 View Figures 110–117 ). Surface of dorsum smooth. Ocularium with only one conspicuous tubercle near the left eye except for the long median spine. Chelicerae of normal shape, not enlarged, slight difference in inner edges of cheliceral finger (Fig. 115 View Figures 110–117 ). Femora of pedipalpi ventrally with six and dorsally with a row of five setiferous tubercles, disto-medially with three setiferous tubercles. Tarsal formula (I-IV): 6(2)/15(2)/7/7.

Ovipositor (Figs 122 View Figures 118–123 , 123 View Figures 118–123 ). Ventral side with four, dorsal side with six setae.

Measurements. Male (female): body 2.27 (2.76) long, 2.08 (1.96) wide at widest portion, scutum 1.95 (1.79) long. Ocularium 0.41 (0.30) long, 0.69 (0.51) wide. Proximal chelicerae 1.24 (0.84) long, 0.47 (0.31) wide; second 2.20 (1.37) long, 0.78 (0.45) wide; distal 0.97 (0.62) long, 0.22 (0.14) wide. Pedipalp claw 0.82 (0.79) long. Penis 1.02 long. Measurements of pedipalp and legs as in Tables 7 View Table 7 and 8 View Table 8 .

Habitat.

The specimens were collected under stones (SMF-CJM7065) and under Acacia bark and rotten stumps (SMF-CJM7063).

Distribution.

Thailand (Loel Province, Nong Khai Province).