Sphecodes geoffrellus ( Kirby, 1802 )

Astafurova, Yulia V. & Proshchalykin, Maxim Yu., 2015, Bees of the genus Sphecodes Latreille 1804 of Siberia, with a key to species (Hymenoptera: Apoidea: Halictidae), Zootaxa 4052 (1), pp. 65-95 : 72-73

Sphecodes geoffrellus ( Kirby, 1802)

( Figs 32, 33 , 64 , 77 , 107 , 115 )

Melitta geoffrella Kirby, 1802: 45 , ♀ (type locality: England).

Sphecodes affinis Hagens, 1882: 224 , ♀, ♂ (type locality: Germany).

Sphecodes fasciatus Hagens, 1882: 224 , ♂ (type locality: Germany); Blüthgen 1923: 508; Cockerell 1937: 4.

Sphecodes rimalis Pérez, 1903: 221 , ♀ (type locality: France).

Sphecodes impunctatus Meyer, 1922: 171 , ♂ (syntypes: 2 ♂, “Jakutsk, 10–18.VII.08 ” [ Russia, Yakutsk], MNHB). Synonymized by Blüthgen 1923: 508 under Sphecodes fasciatus Hagens, 1882 .

Sphecodes silvicola Tsuneki, 1983: 60 , Figs 246–248, ♀ (type locality: Japan).

Sphecodes shirozui Tsuneki, 1983: 53 , Figs 98–104, ♂ (type locality: Japan).

Sphecodes kitamius Tsuneki, 1983: 55 , Figs 105–108 , ♂ (type locality: Japan).

Sphecodes geoffrellus: Warncke 1992: 55 (map); Proshchalykin 2012: 457.

Material examined. Russia. Tomsk Prov.: 1 ♀, Anikino [near Tomsk], 9.VI.1996, OK; Tomsk distr., 1 ♀, Chernyshovka, 14.IX.1996, OK; 4 ♀, Tomsk distr., 41st-kilometer [railway platform], 24.V.1997, OK; 3 ♀, Tomsk, 16.VI.2000, OK; Kemerovo Prov.: 1 ♀, Belovo distr., Bekovo, 26.V.2001, NE; 1 ♀, Leninsk-Kuztetskiy distr., Shebanovo, 14.VII.2004, DS; 1 ♀, Tashtagol distr., Mundybash, 5.VII.2005, DS; 2 ♀, 16 km NWW Kemerovo, on Potentilla humifusa, 16.V.2008, DS; 9 ♀, Belovo distr., Karakan Ridge, on Potentilla chrysantha, 3.VI.2008, DS; 1 ♀, Krapivinskiy distr., bio-station “Azhendarovo”, 15–30.VII.2008, A.Korshunov; 1 ♀, Promyshlenovskiy distr., 4 km W Zhuravlevo, 29.V.2009, DS; 1 ♀, Kemerovo, 7.VI.2009, A.Korshunov; Altaisky Terr.: 1 ♀, 2 ♂, 30 km S Kur’ya, Savvushka, 1.VIII.2007, SB; 1 ♂, 40 km SSE Zmeinogorsk, Novoaltayskoe, 6.VIII.2007, SB; Altai Rebublic: 2 ♂, Chike-Taman Pass, 17.VII.1964, MK; 1 ♂, Artybash, 19.VII.1983, AB; 1♀, Tashanta, 14.VI.1977, V. Rikhter; 1 ♂, 36 km SE Onguday, Terektin Ridge, Bolshoy Yaloman River, 29.VII.1987, YP; 1 ♀, 2 ♂, Yaylyu, Teletskoe Lake, 29.VIII–13.IX.1998, E.Korneeva; 2 ♂, 10 km SW Kuray, 11.VII.2007, SB; 1 ♀, 12–15 km SW Kuray, 12.VII.2007, SB; 1 ♂, Chemal, 19–21.VII.2007, SB; Khakassia Republic: 1 ♀, Shira Lake, 21–24.VI.2011, KT; 2 ♀, 27 km E Shira, 29–30.VI.2014, AL, MP, VL; 2 ♀, 22 km NW Shira, Bely Iyus River, 2.VII.2014, AL, MP, VL; Tyva Republic: 1 ♀, Erzin, 24.V.1975, DK; 3 ♀, Turan, 2–7.VI.1975, DK; 2 ♀, 11 ♂, 30 km W Mugur- Aksa, Maly Khindiktig-Khol [Lake], 2270 m, 8.VIII.1987, YP; 1 ♀, Balgazyn, 29.VII.2009, SB; 4 ♀, 1 ♂, Shuurmak, Shuurmak River, 11–12.VII.2013, MP, VL; 4 ♀, 25 km SE Erzin, Tes-Khem River, 14–15.VII.1014, AL, MP, VL; 1 ♀, 31 km NEE Erzin, Erzin River, 16, 18.VII.2014, AL, MP, VL; 1 ♀, 6 km SE Bay-Khaak, Sosnovka, 20–21.VII.2014, AL, MP, VL; Krasnoyarsk Terr.: 1 ♀, Sukhoy Buuzim, 45 km ENE Krasnoyarsk, 11.VI.1903, Salstrem; 1 ♀, “Stolby” Nature Reserve, Enisey River, 19.VI.2011, KT; 1 ♀, Bol’shoy Arey River [near Krasnoyarsk], 12.VI.2014, EA; 1 ♀, Solontsy, 30.VI.2013, EA; 3 ♀, Magansk, 15.VI.2014, EA; Irkutsk Prov.: 2 ♀, Padun [near Bratsk] no date, A. Chekanovskiy; 7 ♀, 3 ♂, Irkutsk, no date [1892–1898], V.Yakovlev; 1 ♀, Tibel’ti, 35 km W Baikal Lake, 3.VI.1970, DK; 1 ♀, 1 ♂, 15 km E Ust’-Ordynskiy, 31.VII.1994, AL; Buryatia Republic: 1 ♀, Romanovka, 4.VII.1983, D.Shcherbakov; 1 ♂, Naushki, Selenga River, 30.VII.2007, AL, MP, VL; 1 ♂, Gusinoe Lake, Baraty, 25.VII.2007, AL, MP, VL; 8 ♀, Selenduma, 19–24.V.2008, MP; 1 ♀, Bilyutay, 23.V.2008, MP; 1 ♀, Ust’-Kiran, Chikoy River, 27–28.V.2008, MP; 1 ♀, Kurumkan distr., NR “Jerginskiy”, 4– 5.VII.2013, S.Rudykh; Yakutia: 1 ♀, 5 ♂, Yakutsk, 24.VII–1.VIII.1927, Moskvin; 1 ♀, Bestyakh, 30.V.1912, Naumov; 1 ♀, 25 km S Yakutsk, Lena River, 27.VI.1970, DK; 3 ♀, Artyk, Nera River, 14.VII.1974, YP; 8 ♀, Balagannakh, 30 km ESE Ust’-Nera, 4–15.VII.1974, YP; 1 ♀, Indigirka, 15 km S Tebyulyakh, 9.VII.1974, YP; 3 ♀, Khaptagay, Lena River, 30 km SSE Yakutsk, 28–30.VI.1974, YP; 1 ♀, Abaga- 1, 10 km S Olekminskiy, 5.VIII.1974, EN; 2 ♀, Mikhaylovka, 60 km N Amga, 25.VII.1985, Bagachanova, Kaymuk; 1 ♂, Tyungyulyu, 50 km ENE Yakutsk, 11.VIII.1990, Kashirtseva; 1 ♀, Lena, River, mouth of Labyya River, 1.VIII.2000, ND; Zabaikalskiy Terr.: 3 ♀, Bylyra, Kira River, 19.VI.1975, DK; 2 ♂, Kalga, 13.VII.1975, DK; 1 ♂, 7 km N Priargunsk, 25.VII.1975, DK.

Distribution. Russia: Tomsk Prov. ( Warncke 1992), *Kemerovo Prov., *Altaisky Terr., *Altai Rebublic, *Khakassia Republic, *Tyva Republic, *Krasnoyarsk Terr., Irkutsk Prov. ( Blüthgen 1923), *Buryatia Republic, Yakutia ( Meyer 1922 as Sphecodes impunctatus ), *Zabaikalskiy Terr., Far East, European part ( Osytshnjuk et al. 1978); Japan, Mongolia, N Kazakhstan ( Popov 1934); Europe (north to 66°), Turkey, Near East, North Africa.

Figures

FIGURE 115. Distribution of Sphecodes species in Siberia: a— S. albilabris (Fabricius), S. alternatus Smith; b— S. crassus Thomson, S. cristatus Hagens; c— S. ephippius (Linné), S. ferruginatus Hagens; d— S. gibbus (Linnaeus); S. geoffrellus (Kirby); e— S. longulus Hagens, S. hyalinatus Hagens; f— S. miniatus Hagens, S. monilicornis (Kirby).

FIGURES 24 – 45. Males, gonostylus, dorsal and lateral views: 24, 25 — Sphecodes albilabris (Fabricius); 26, 27 — S. cristatus Hagens; 28, 29 — S. crassus Thomson; 30, 31 — S. ferruginatus Hagens; 32, 33 — S. geoffrellus (Kirby); 34, 35 — S. ephippius (Linné); 36, 37 — S. hyalinatus Hagens; 38, 39 — S. kozlovi Astafurova & Proshchalykin; 40, 41 — S. longulus Hagens; 42, 43 — S. miniatus Hagens; 44, 45 — S. monilicornis (Kirby). Dark indicate thickened areas. Scale bar: 0.5 mm.

FIGURES 62 – 68. Males, head: 62 – 64 — frontal view; 65 – 68 — dorsal view; 62 — Sphecodes pinguiculus Pérez; 63 — S. schwarzi, sp. nov.; 64 — S. geoffrellus (Kirby); 65 — S. cristatus Hagens; 66 — S. nippon Meyer; 67 — S. reticulatus Thomson; 68 — S. longulus Hagens.

FIGURES 77 – 88. Females, head: 77 – 84 — frontal view; 85 – 88 — dorsal view; 77 — Sphecodes geoffrellus (Kirby); 78 — S. monilicornis (Kirby); 79 — S. longulus Hagens; 80 — S. crassus Thomson; 81 — S. puncticeps Thomson; 82 — S. pellucidus Smith; 83 — S. ephippius (Linné); 84, 85 — S. gibbus (Linnaeus); 86 — S. reticulatus Thomson; 87 — S. scabricollis Wesmael; 88 — S. pinguiculus Pérez.

FIGURES 105 – 114. Females: 105, 106 — pronotum; 107 – 110 — head, frontal view; 111, 112 — hind wing; Males: 113, 114 — genitalia, dorsal view; 105, 114 — Sphecodes ferruginatus Hagens; 106 — S. miniatus Hagens; 107 — S. geoffrellus (Kirby); 108 — S. crassus Thomson; 109, 111 — S. ephippius (Linné); 110 — S. pellucidus Smith; 112 — S. nippon Meyer; 113 — S. spinulosus Hagens.