Bembidion lugubre, LeConte, 1857

Maddison, David & Swanson, Alexander, 2010, A preliminary characterization of Bembidion perspicuum LeConte, with a reclassification of related species (Coleoptera, Carabidae) north of México, ZooKeys 43 (43), pp. 15-31 : 28-29

B. lugubre LeConte, 1857

Bembidium lugubre LeConte, 1857:6 , described as a “variety” of B. mexicanum Dejean. Type locality “ Valley of the Rio Grande ”. Holotype female in MCZ (type number 5511); external structure and spermatheca examined.

Bembidium sallaei Bates, 1882:148 . Type locality not specified, with localities specified in several states of México as well as Belize and Guatemala. Lectotype male, designated by Erwin (1982), in MNHN; external structure and aedeagus examined.

Bembidion canonicum Casey, 1918:61 . Type locality Arizona. Holotype male in USNM (type number 36920); external structure and aedeagus examined.

Bembidion retectum Casey, 1918:61 . Type locality St George, Utah. Lectotype male, designated by Erwin (1984), in USNM (type number 36923); external structure and aedeagus examined.

This species is known from Texas, New Mexico, Arizona, Utah, and California south at least to Oaxaca, México. In the United States specimens of this species tend to be paler than those of B. mexicanum , with some specimens having a pale elytral apex, and with more notable punctures at the base of the pronotum. Male genitalia, female spermathecae, and DNA sequences are distinctive.