Clivina agona Putzeys, 1866

Balkenohl, Michael, 2021, Belgian Journal of Entomology, Belgian Journal of Entomology 115, pp. 1-83 : 24-26

publication ID 10.5281/zenodo.5645953


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Clivina agona Putzeys, 1866

stat. restit.

Clivina agona Putzeys, 1866 View in CoL stat. restit.

( Figs 13 View Figs 9–15 , 20 View Fig , 40 View Fig , 60 View Fig , 78 View Fig , 87 View Fig , 107 View Fig )

Clivina agona PUTZEYS, 1866: 131 View in CoL ; CSIKI, 1927: 497;

Clivina agona PUTZEYS, 1866 View in CoL : ANDREWES 1926: 377 ( C. parryi Putzeys View in CoL );

Clivina javanica var. agona PUTZEYS, 1866 View in CoL : ANDREWES 1930b: 113;

Clivina javanica agona PUTZEYS, 1866 View in CoL : LORENZ, 1998: 132, 2005: 143; BALKENOHL, 2001: 15.

TYPE MATERIAL. Holotype: • ♂, mounting card with small pin, with labels and data: cream white, handwritten in black ink “ Siam Casteln ” / yellow, handwritten in black ink “ Cliv. agona Putz type Siam y Castelnau ” / small, grey brown “5” / yellow, handwritten in black ink “ agona Putz. ” / white, printed “Chevrolat Carabidae . Fr. V.d Poll. Pres. 1909, E.B. Poulton.” / white, handwritten in black ink and black printed “ Clivina castanea, Westw. = parryi, Putz. Teste 1918 . H.E. Andrewes.” / white, large, black framed, black printed and handwritten in black ink “TYPE COL: 811 Clivina agona Putzeys HOPE DEPT.OXFORD ” ( Fig. 13 View Figs 9–15 ) ( OUMNH) .

REMARKS. The following parts are missing: Right front leg with tarsomeres two to five; left front leg with all tarsomeres; right intermediate leg tarsomeres two to five; right hind leg completely missing. The right elytron shows a hole where the small original pin was inserted.

PUTZEYS (1866) based his description on one specimen.

ADDITIONAL MATERIAL. CAMBODIA. • 1 ♀, Coll. I.R.Sc.N.B. CAMBODIA Banteay M. Prov. Ang Trapeang Thmor W.S. Light Trap Office Int. Crane F., leg. Var & Hagebaert ( RBINS) ; • 19 specs., Coll. I.R.Sc.N.B. CAMBODIA Prek Toal (Tonle Sap Lake) 27.v.2003 – Light trap, leg. J. Constant, K. Smets & P. Grootaert ( RBINS) ; • 10 specs., same data but 06.vii.2005 day catch, leg. I. Var ( RBINS) ; • 10 ♂♂, 34 ♀♀, 14 specs., Coll. I.R.Sc.N.B. CAMBODIA Siem Reap 22.v.2003 and 25.v.2003, Light trap, leg. J. Constant & K. Smets ( RBINS) ; • 1 spec., CAMBODIA: Siem Reap, 13°21’N, 103°51’ Ekilled by halogen light reflector, 7–11.xi.2002, 20 m a.s.l. P. Kočarek leg. / CLIVINA (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2016 ( NMHP) GoogleMaps ; • 5 spec., CAMBODGE / Collection P. Dupuis (one with damaged abdomen) ( RBINS) ; • 1 spec., CAMBODGE PNOMH PËNH VITALIS / Collection P. Dupuis ( RBINS) ;

THAILAND. • 1 ♂, 1 ♀, THAILAND, S 40 km NE OF BETONG, LAKE BANG LANG (KG.TO) 12– 2007, 350 m Z. LINEK LGT. ( CADW); • 1 historic spec., Siam Gutti / Clivina agona Putz. dét. J. Putzeys / Soc. Ent. Belg. Coll. PUTZEYS / Clivina castanea Westw. H.E. Andrewes det. 1928 ( RBINS) ; • 1 spec., SAIGON (ville) 7. viii.1903 Cap. Fouquet. / Clivina castanea Westw. = Parryi Putz. H. E. Andrewes det. ( ETHZ) .

DIAGNOSIS. Amedium sized species with smooth lateral margin of the pronotum and elytron, and isodiametric reticulation on interval eight of the elytron. Distinguished from the other species with isodiametric reticulation on interval eight of the elytron by the group of distinct punctures on each side of the pronotum. The only other species with a group of distinct punctures laterally on the pronotum is C. trachys . However, that species shows two tubercles at the base of the pronotum and the lobe of the mentum is double pointed picked, and all setigerous punctures of the third interval of the elytron are approaching the third stria.

REDESCRIPTION. Measurements. Holotype: Body length 8.53 mm, width 2.63 mm; ratio length/width of pronotum 0.91; ratio length/width of elytra 1.78. Other material: Body length 8.3–8.94 mm (: = 8.59 mm *), width 2.32–2.64 mm (: = 2.56 mm *), ratio length/width of pronotum 0.9–0.93 (: = 0.92*), ratiolength/widthofelytra 1.78–1.86 (: = 1.84*); (*n = 10).

Colour: Glossy. Piceous. Antenna and tarsomere fuscous.

Head: Athird narrower than pronotum. Clypeus excised anteriorly; wing nearly as protruding as clypeus, flattened, separated from clypeus by distinct notch; supraantennal plate convex, reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by sharp notch. Clypeus square, moderately convex in posterior part, separated from frons by moderately deep furrow. Frons convex, with moderately big pore at middle, with posteriorly increasing blunt lateral carina at posterior eye-level. Clypeus and frons smooth, separated from wing and supraantennal plate by deep moderately wide furrow continuing posteriorly as wide supraorbital furrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as step, with row of small punctures. Eye hemispherical projecting. Gena short. Antenna reaching just over posterior angle of pronotum, antennomeres four to ten elongate (ratio L/W about 2.04). Labrum slightly trilobed, with isodiametric reticulation, with transverse carina, seven setose. Mentum ( Fig. 40 View Fig ) with isodiametrically to longitudinally reticulated surface, shape of lobe triangular with anterior angle acute, rounded at tip, anterior margin slightly convex, carinate laterally and anteriorly, with distinct carina at middle, median tooth wide, distinctly less protruding anteriorly as lobe, obtusely angled anteriorly.

Pronotum ( Fig. 60 View Fig ): Disk moderately convex in lateral and distinctly convex frontal view. Wider as long. Reflexed lateral margin smooth, convex, widest in posterior third; anterior angle obtusely rounded off, not projecting; posterior angle marked by distinctly sharp small tooth. Lateral channel reticulated, with some indistinct foveae. Median line narrow, line-like, somewhat wider in posterior third, narrower anteriorly, anterior transverse line striate-punctate, median line joining and surpassing anterior transverse line, joining basal constriction. Disk laterally with group of distinct punctures of moderate size, surface with some transverse and a few longitudinal wrinkles, with band of isodiametric reticulation towards lateral channel and in basal third, with distinct transverse rugae at basal declivity.

Elytron: Disk flattened in anterior half in lateral view, convex on frontal view. Less than twice as long as wide, slightly dilated in apical half with maximum width behind middle. Reflexed lateral margin smooth. Scutellar striole long; setigerous tubercle at base of first stria, with distinct tubercle at base of third interval. Striae moderately deep, punctuate-striate, one to three free at base, four to seven joining at humerus, one joining with seven at apex, two, three and four ending free at apex, five and six joining indistinctly at apex. Intervals moderately convex, eight with short carina at humerus and longer one apically. Third interval with four setigerous punctures, first approaching second stria near base, the other three approaching third stria. Surface of all intervals glossy, with isodiametric reticulation on interval eight.

Hind wings: Fully developed.

Lower surface: Proepisternum covered completely with distinct isodiametric reticulation and with distinct coriaceate-like wrinkles ( Fig. 78 View Fig ). Terminal sternite of abdomen at apex with flat transverse sulcus.

Legs: Legs covered with distinct reticulation. Protibia with three spines of moderate length, nearly not sulcate dorsally, movable spur slightly arcuate in basal three quarters, distinctly arcuate in apical quarter.

Male genitalia ( Fig. 87 View Fig ): Median lobe slender, of same breadth in nearly whole length, moderately arcuate, distinctly arcuate at base, ventral rim distinct, apex developed as triangular slightly asymmetrical spatula of moderate length. Endophallus densely folded, apically with longitudinal group numerous bristles. Ventral paramere moderately wide, moderately arcuate, lateral apophysis stout, apex long tapered, with four long setae. Dorsal paramere slender, regularly arcuate, at apex with four long setae.

Female coxostylus and laterotergite ( Fig. 107 View Fig ): Coxostylus moderately broadened at base, dorsally with ten long and slender and one distinctly elongated and robust nematiform setae in basal half, ventrally with numerous smaller setae, with two SSOs apically. Laterotergite with four long and two small setae basally and one isolated seta towards coxostylus.

VARIATION. The margin of the clypeus is more or less distinctly reflexed. It is not reflexed in the holotype but developed to a slight degree in some of the other specimens. The reflexed lateral margin of the pronotum is slightly undulated in some of the specimens. The group of punctures present laterally on the posterior half of the disk of the pronotum is always clearly visible. The character is distinct in the holotype and in most of the other specimens. However, the number of punctures varies, in occasional cases there are only around six punctures on each side.

DISTRIBUTION. Known from several places in the centre of Cambodia, from Thailand and from Vietnam.

REMARK. ANDREWES (1919) considered C. agona as a distinct species. Later, he considered C. agona as slight variety and synonymized C. agona with C. castanea ( ANDREWES, 1926) . In 1929, the species was not considered at all ( ANDREWES, 1929a), and in the Catalogue of Indian Insects C. agona was treated as variation of “ C. javanica Bouchard, 1903 ” ( ANDREWES, 1930b). In reality, BOUCHARD (1903) refers in his contribution to C. javanica Putzeys, 1846 . The confusion was increased by a type writing error of BOUCHARD (1903: 169) who quoted the page of PUTZEYS description (1846) as “529” instead of 592. As PUTZEYS (1866) already indicated, C. javanica Putzeys, 1846 is similar to C. advena Putzeys, 1866 . Both species are members of the Clivina lobata -species group.


United Kingdom, Oxford, University Museum of Natural History






Switzerland, Zurich, Erdgenoessische Technische Hochschule-Zentrum


Royal Belgian Institute of Natural Sciences


Eidgenoessische Technische Hochschule-Zentrum














Clivina agona Putzeys, 1866

Balkenohl, Michael 2021

Clivina javanica agona

LORENZ W. 2005: 143
BALKENOHL M. 2001: 15
LORENZ W. 1998: 132

Clivina javanica var. agona

ANDREWES H. E. 1930: 113

Clivina agona PUTZEYS, 1866

ANDREWES H. E. 1926: 377

Clivina agona PUTZEYS, 1866: 131

CSIKI E. 1927: 497
PUTZEYS J. A. A. H. 1866: 131
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