Clivina plurisetofaria, Balkenohl, 2021

Balkenohl, Michael, 2021, Belgian Journal of Entomology, Belgian Journal of Entomology 115, pp. 1-83 : 32-34

publication ID 10.5281/zenodo.5645953


persistent identifier

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scientific name

Clivina plurisetofaria

sp. nov.

Clivina plurisetofaria View in CoL sp. nov.

( Figs 8 View Fig , 24 View Fig , 44 View Fig , 64 View Fig , 91 View Fig , 110 View Fig )

ETYMOLOGY. The name plurisetofaria refers to the female genitalia and is composed from the Latin adverb ‘plurifariam’ = at many places, and the Latin female noun ‘seta’ = long bristle. ♂

TYPE MATERIAL. Holotype: • ♂, with labels and data: yellow, black printed “S- THAILAND, Khao Sok rainforest, 38 km E Takua Pa , 21.xi.1996, leg. J. REJSEK ” ( SMNS) .

REMARKS. The following parts are missing: left front claw, tarsomere four and five of left hind leg.

Paratypes: THAILAND. • 1 ♂, NW THAILAND, Mae Hong Son, 29.iv.1982, J. HORAK legit ( CADW) ; • 4 ♂♂, NW THAILAND, Mae Hong Son Huai Sua Tao , 11–17.v.1992, Strnad Jan leg. ( CADW) ; • 2 ♂♂, 1 ♀, 1 spec., same data as holotype ( SMNS / CMBB) ; • 1 spec. W-THAILAND, Sangkhlaburi , xi.1994, leg. J. REJSEK ( SMNS) ; • 1 ♂, THAI, 29.iv.1993, PAI City Pacholatko & Dembicky leg. ( CADW) ; • 14 ♂♂, 15 ♀♀, S-Thailand, Betong Gunung Cang dun vill Yala dist., 25.iii.–22.iv.1993, J. Strnad leg. ( CADW) ; • 4 ♂♂, 7 ♀♀, same data but J. Horák leg. ( CADW) ; • 1 ♂, 2 ♀♀, THAILAND, S. 40 km NE OF BETONG , LAKE BANG LANG (KG.TO), 12– 2007, 350 m Z. LINEK LGT. ( CADW) ; • 1 spec., Siam Bgk. / Clivina lata Putz. dét. J. Putzeys / Soc.Ent. Belg. Coll. PUTZEYS ( RBINS) .

VIETNAM. • 1 spec., SAIGON (ville) 7.viii.1903 Cap. Fouquet / Clivina parryi Putz. ( ETHZ) . CAMBODIA. • 1 spec., I.R.Sc.N.B, Cambodia, 8 km north of Sre Noi (road to Along Vaeng), Light trap 29.v.2003, J. Constant & K. Smets ( RBINS) .

MALAYSIA. • 13 specs., MALAYSIA W KELANTAN 30 km NW of Gua Musang Ulu Lalat Mt. 800–1000 m KAMPONG SUNGAI OM 25.v.–, Petr Cechovsky leg. ( CADW) ; • 1 spec., MALAYSIA (Phang State) Kuala Tahan 29.x.1990 Guillaume de Rougemont / Clivina castanea Westwood, 1837 , P. Bulirsch det. 2013 (CWGP); • 1 spec., MALAYSIA Phang Taman Negara. 1–13.iii.1984 / at light / L.Jessop. B.M.1984-230 ( BMNH) ; • 1 spec., MALAYSIA (Kelantan State) 60 km NE Tanah Rata Tanah Kerajaan 1000 m (at light) 12– 30.iv.2007, P. Čechowsky ( CWGP) .

DIAGNOSIS. Amedium sized species with smooth lateral margin of the pronotum and elytron, and isodiametric reticulation on all intervals of the elytron. Distinguished from all other species by the reticulation of the disk of the pronotum and the unusual number and pattern of setae on the female coxostylus and laterotergite.

DESCRIPTION. Measurements. Holotype: Body length 8.54 mm, width 2.57 mm; ratio length/width of pronotum 0.92; ratio length/width of elytra 1.87. Other material: Body length 7.9–8.9 mm (: = 8.31 mm *), width 2.39–2.67 mm (: = 2.51 mm *), ratiolength/width of pronotum 0.89–0.93 (: = 0.91*), ratiolength/widthofelytra 1.84–1.9 (: = 1.88*); (* n = 10).

Colour: Semi-opaque, shiny. Piceous. Intermediate and hind leg dark fuscous, antenna, mouthparts and tarsomeres fuscous.

Head: More than a third narrower than pronotum. Clypeus slightly excised anteriorly, with lateral angle rounded; wing less protruding as clypeus, hollowed out anteriorly, separated from clypeus by distinct notch; supraantennal plate convex; clypeus and supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by obtuseangled notch. Clypeus transverse, slightly convex, separated from frons by distinct but not deep furrow. Frons moderately convex, with pore at middle, converging lateral carina at posterior eye-level prolonged anteriorly. Clypeus and frons with scattered microscopic punctures, separated from wing and supraantennal plate by deep and wide furrow continuing posteriorly as wide supraorbital furrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as step, with irregular punctures. Eye laterally projecting, somewhat elongated in dorsal view. Gena small. Antenna relatively short, just reaching posterior setigerous puncture of pronotum, antennomeres four to ten elongate (ratio L/ W 1.94). Labrum obtusely pointed to convex anteriorly, with indistinct reticulation, seven setose. Mentum ( Fig. 44 View Fig ) with distinct isodiametrically reticulated surface, shape of lobe like a skewed trapezium with conspicuously acute antero-lateral angle (angle around 80°) and obtuse medial angle, reflexed margined, anterior margin askew, with epilobe laterally indistinct and becoming slightly wider towards middle, with distinctly long keel at middle, median tooth wide, anteriorly nearly as protruding as lobe, obtusely angled anteriorly (angle around 140°).

Pronotum ( Fig. 64 View Fig ): Disk regularly convex in lateral view and frontal view. Outline square to globose, wider than long. Reflexed lateral margin smooth, regularly and slightly convex in anterior half but converging anteriorly, more convex in posterior third, widest at posterior third; anterior angle obtuse, slightly projecting; posterior angle developed as distinct tooth. Lateral channel smooth, of same width between setigerous punctures. Median line narrow, fine anteriorly and posteriorly; anterior transverse line sharp, wider as median line, joining and surpassing median line. Surface with indistinct transverse wrinkles, completely covered or with patterns of isodiametric reticulation, with scattered microscopic punctures, base with few transverse rugae at declivity.

Elytron: Disk flattened in anterior half and regularly convex to apex in lateral view, regularly convex in frontal view. Distinctly less than twice as long as wide, nearly parallel at middle, maximum width at middle. Reflexed lateral margin smooth. Scutellar striole distinct. Base of elytron with setigerous tubercle at base of first stria, with a small tubercle at base of third interval. Striae relatively deep, punctuate-striate, one to three free at base, four to seven joining at humerus, one joining with seven at apex, two to four ending free, and five and six joining at apex. Intervals moderately convex, eights with carina at humerus and apex, Third interval with four setigerous punctures, first close to base and situated at the second stria, the other three approaching third stria. Surface of intervals semi-opaque, covered with isodiametric reticulation, reticulation becoming more evident laterally.

Hind wings: Fully developed.

Lower surface: Proepisternum covered with distinct isodiametric reticulation, with some transverse wrinkles of moderate size. Terminal sternite of abdomen developed apically as deep narrow transverse fold.

Legs: Legs covered with isodiametric reticulation. Protibia with two longer and one smaller spines, slightly sulcate and carinate anterior-dorsally, movable spur apically stronger arcuate.

Male genitalia ( Fig. 91 View Fig ): Median lobe stout, robust, regularly arcuate, of same diameter in the course of situation, apex moderately long, tapered nearly parallel in lateral view, spatula slightly distorted, apex regularly rounded, ventral rim distinct nearly up to base, base with small apophysis. Endophallus with band of small bristles at both lateral sides. Ventral paramere distinctly widened, wider at middle, lateral apophysis evident, the other one moderately developed; apex short, arcuate, with two long and robust and two small setae at apex. Dorsal paramere slender, arcuate in apical third, with indistinct apophyses, with three long setae at apex and one small seta preapically.

Female coxostylus and laterotergite ( Fig. 110 View Fig ): Coxostylus of moderate size, apical third flattened with sharp margins, setae arranged dorsally conspicuously regular in two rows, dorsally with eleven long to medium sized and two distinctly elongated and robust nematiform setae in basal half, ventrally with around twenty smaller setae, no SSO visible (500×). Laterotergite with eight medium to long setae basally and two isolatedsetae towards coxostylus.

VARIATION. The pore on the frons of the head is more or less evidently developed. In some specimens striae five and six on the elytron are joining apically at one side only. There are also few specimens where striae three and four joining at apex.

DISTRIBUTION. Known from Thailand, Vietnam, Cambodia, and Malaysia.


Germany, Stuttgart, Staatliches Museum fuer Naturkunde






Switzerland, Zurich, Erdgenoessische Technische Hochschule-Zentrum


United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]




Staatliches Museum fuer Naturkund Stuttgart


Royal Belgian Institute of Natural Sciences













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