Clivina germanni, Balkenohl, 2021

Balkenohl, Michael, 2021, Belgian Journal of Entomology, Belgian Journal of Entomology 115, pp. 1-83 : 30-32

publication ID 10.5281/zenodo.5645953


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scientific name

Clivina germanni

sp. nov.

Clivina germanni View in CoL sp. nov.

( Figs 23 View Fig , 43 View Fig , 63 View Fig , 90 View Fig , 109 View Fig )

ETYMOLOGY. The name is a patronym in honour of Dr. Christoph Germann (NHMB) who provided me full access to the Collection Georg Frey, Basel, and who helped me to localize a lot of material of this contribution.

TYPE MATERIAL. Holotype: • ♂, with labels and data: beige, black printed “ Kina Balu Borneo ” / white, handwritten with black ink “ Clivina castanea Westw. ” ( NHMB Frey) .

REMARKS. The following parts are missing: Left front tarsomeres three to five, tarsomeres of left hind leg damaged.

Paratypes: BORNEO. • 1 ♂, 4 ♀♀ ,, same data as holotype but without determination label ( NHMB Frey/CMBB) ; • 1 ♀, with labels and data: white, black printed 19.iii.1987 Borneo Sarawak Belaga Distr. Belage leg. Hiermeler ( CMBB) ; • 1 spec., Borneo, Malaysia, Sarawak Kubah NP nr. Matang Wildlife Centre , 19–22.ix.2008 R. Grimm ( CMBM) ; • 2 ♂♂, Sarawak, Kapit dist. Sebong, Baleh riv. 9–21.iii.1994 J. Horák leg. ( CADW) ; • 2 specs., Borneo c. Coll. H. Müller ( CADW) ; • 7 specs., Coll. I.R.Sc.N.B. BORNEO: ca 400 km NNE Pontianak , Putussibau-N env., 0°53’N, 112°56’E, 2-10-I-2009 I.G.31;412, leg. A. Napolov ( RBINS) GoogleMaps ; • 1 spec., Borneo, Sarawak, Kubah NP, Matang Wildlife Centre env., 50 m, 8–11.iv.2016, R. Grimm ( CMBM) ; • 1 specs., Borneo, Sabah Tambunan , 500 m, 30.iii.2015, R. Grimm ( CMBM) ; • 2 specs., same data but 19–22.iii.2015 ( CMBM) ; • 2 specs., Borneo, Sarawak, Kota Kinabalu NP, 500 m Tambunan , 4–8.ii.2014, R. Grimm ( CMBM) ; • 1 spec., Borneo, Malaysia, Sabah Sepilok , 100 m 31.iii–1.iv.2015, R. Grimm ( CMBM) ; • 2 ♂♂, 1 spec., SARAWAK (Borneo), ca. 25 km E Kapit . iii.1994, Kodada leg. ( NHMW) ; • 3 specs., MALAYSIA, Sarawak Mulu NP , 3.iii.1993 leg. M. Jäch (19) ( NHMW) .

SUMATRA. 2 ♂♂, 5 ♀♀, with labels and data: white, black printed “I. W. Sumatra Harav vall. II-III. 1991 ” ( CMBB; CADW); 1 spec., Sumatra 19 I 85 Lubu- Sampir / C. castanea Westw. ssp. nov. Kult 1946 ( NHMP) .

NIAS. 1 spec., INDONESIA; Nias Ostküste: Lawalo 26.9.1979 ERBER leg. Lichtfang / BM 1984 2 ( BMNH) .

REMARK. In few of the specimens the body surface shows numerous microscratches. These scratches camouflage the fine natural reticulation on the elytron and pronotum.

DIAGNOSIS. Amedium sized species with smooth lateral margin of the pronotum and elytron, isodiametric reticulation on all intervals of the elytron, and with one tubercle at the base of interval three. Mainly distinguished from C. lata by the smaller size, the stout askew trapezium-like lobe of the mentum, and by the female laterotergite which shows 8 setae with one of them isolated. Distinguished mainly from C. plurisetofaria sp. nov. by the slender and anterolaterally acute lobe of the mentum, and be the female laterotergite which shows then setae with two of them isolated.

DESCRIPTION. Measurements. Holotype: Body length 8.26 mm, width 2.48 mm; ratio length/width of pronotum 0.95; ratio length/width of elytra 1.84. Other material: Body length 7.01–8.51 mm (: = 8.12 mm *), width 2.02–2.69 mm (: = 2.45 mm *), ratiolength/width of pronotum 0.89–0.93 (: = 0.92*), ratiolength/widthofelytra 1.81–1.91 (: = 1.84*); (*n = 10).

Colour: Semi-opaque, shiny. Piceous. Intermediate and hind leg dark fuscous, antenna and tarsomeres fuscous.

Head: Athird narrower than pronotum. Clypeus slightly excised anteriorly, with lateral angle rounded; wing slightly less protruding as clypeus, flattened and slightly hollowed out anteriorly, separated from clypeus by distinct notch; supraantennal plate convex; clypeus and supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by obtuse-angled notch. Clypeus transverse, moderately convex posteriorly, separated from frons by distinct but not deep furrow. Frons moderately convex, with small flat pore at middle, with converging lateral carina at posterior eye-level. Clypeus and frons with some fine reticulation, separated from wing and supraantennal plate by deep and wide furrow continuing posteriorly aswide supraorbital furrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as step, with irregular punctures. Eye laterally projecting. Gena indistinct. Antenna relatively short, just reaching posterior setigerous puncture of pronotum, antennomeres four to ten of moderate length (ratio L/W around 1.8). Labrum obtusely pointed anteriorly, isodiametrically reticulated, seven setose. Mentum ( Fig. 43 View Fig ) with distinct isodiametrically reticulated surface, shape of lobe like a skewed trapezium with acute antero-lateral and obtuse medial angle, reflexed margined, anterior margin straight, askew, with short and somewhat indistinct carina at middle, median tooth wide, anteriorly as protruding as lobe, obtusely angled anteriorly.

Pronotum ( Fig. 63 View Fig ): Disk regularly convex in lateral view and frontal view. Outline square to globose, wider than long. Reflexed lateral margin smooth, regularly and slightly convex in anterior half, more convex in posterior third, widest at posterior third; anterior angle distinct, slightly projecting; posterior angle developed as small but distinct tooth. Lateral channel smooth, of same width between setigerous punctures. Median line narrow, fine; anterior transverse line sharp, wider as median line, joining and surpassing median line. Surface with indistinct transverse wrinkles, with isodiametric reticulation, base with few slight transverse rugae at declivity.

Elytron: Disk flattened in anterior third and regularly convex to apex in lateral view, regularly convex in frontal view. Distinctly less than twice as long as wide, slightly dilated in apical half, maximum width behind middle. Reflexed lateral margin smooth. Scutellar striole distinct. Base of elytron with setigerous tubercle at base of first stria, with a small tubercle at base of third interval. Striae relatively deep, punctuate-striate, one to three free at base, four to seven joining at humerus, one just not joining with seven at apex, two to six ending free at apex. Intervals moderately convex, eights with carina at humerus and apex, Third interval with four setigerous punctures, first close to base and situated at the second stria, the other three approaching third stria. Surface of intervals semiopaque, covered with isodiametric reticulation.

Hind wings: Fully developed.

Lower surface: Proepisternum covered with distinct isodiametric reticulation with the hexagonal fields raised, with few not deep transverse wrinkles. Terminal sternite of abdomen developed apically as deep narrow transverse fold.

Legs: Legs covered with isodiametric reticulation. Protibia with two longer and one smaller spines, slightly sulcate and carinate anterior-dorsally, movable spur apically stronger arcuate.

Male genitalia ( Fig. 90 View Fig ): Median lobe robust, regularly arcuate, apex moderately long, tapered continuously in lateral view, slightly bisinuate and turned dorsally, spatulate, tip straight but asymmetrical rounded, ventral rim distinct nearly up to base, base with apophysis. Endophallus strongly folded, spines not visible. Ventral paramere moderately wide, wider at middle, lateral apophysis distinctly elongated, the other one distinct, apically arcuate, with two long and robust setae at apex, with three moderately long and one short seta preapically. Dorsal paramere slender, wider basally, regularly arcuate, with indistinct apophyses, with four long setae at apex, with two short seta preapically.

Female coxostylus and laterotergite ( Fig. 109 View Fig ): Coxostylus slender, dorsally with six long to medium sized and two distinctly elongated and robust nematiform setae in basal half, ventrally with around nine smaller setae, no SSO visible (500 x). Laterotergite with six long and three small setae basally and one isolated seta towards coxostylus.

VARIATION. In specimens from Borneo striae five and six of the elytron joining apically in half of the cases.

DISTRIBUTION. Known from Borneo (Kina Balu) and the West of Sumatra.


Switzerland, Basel, Naturhistorisches Museum








Austria, Wien, Naturhistorisches Museum Wien




United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]


Natural History Museum Bucharest


Royal Belgian Institute of Natural Sciences


Naturhistorisches Museum, Wien













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