Tokyosoma hallum, Mikhaljova & Korsós, 2003

Mikhaljova, E. V. & Korsós, Z., 2003, Millipedes (Diplopoda) From Korea, The Russian Far East, And China In The Collection Of The Hungarian Natural History Museum, Acta Zoologica Academiae Scientiarum Hungaricae 49 (3), pp. 215-242 : 217-219

Tokyosoma hallum sp. n.

( Figs 1–5 )

1 m (holotype), 3 mm, 5 ff (paratypes) – South Korea, Cheju Prov. , Halla-san National Park, ca . 1300 m, 126°00’E, 33°15’N, border zone between the mixed deciduous forest and the mixed evergreen (pine) forest, litter, from beneath stones and trunks, No. 1657., 30.Oct.1993., leg. PEREGOVITS L. & RONKAY L. Male holotype, 2 mm, 3 ff paratypes are deposited in the HNHM , 1 m, 1 f paratypes in ZMUM , 1 f paratype in IBSV .

Etymology: The specific epithet refers to the locus typicus (Halla-san National Park).

Description: Male. Length 12–12.5 mm, width 1.7–1.8 mm with and 1.3–1.4 mm without paraterga.

Coloration: Body in alcohol light beige. Antennae brown, eyes black, legs brownish, increasingly darker distad.

Body with 29 segments. Head setose, vertigial suture visible. Genae strongly convex, setose, labrum usual. Eye patches triangular, each composed of 22–23 ocelli. Antennae long and slender, antennomere 3 longest. Collum semicircular, with 3+3 macrochaetae. Head with genae subequal in width to somite 3. Gnathochilarium usual. Paraterga beginning from somite 2, well-developed on somites 7–25, like swellings on somite 26, missing on somites 27–29. Macrochaetae in a transverse row on somites 27–28, like a very extended triangle on somites 26, like an extended triangle on preceding somites. Macrochaetae long, pointed apically. Caudolateral and medial macrochaetae subequal in length, anterolateral ones shortest. Axial suture well-developed.

Legs long and slender, each claw with both two minute accessory claws dorsally and spinicle (= filament) ventrally at base. Legpairs 1 and 2 typically reduced, each with shortest claw spinicle. Legs 3–7 somewhat enlarged toward gonopods. Tarsal papillae absent. Legs 10 and 11 with coxal glands; both coxa 10 and one 11 with ventral small outgrowth ( Figs 1–2 ); trochanter 11 with stickshaped caudal process.

Anal valves obtusangular at caudal edge in lateral view, each with some marginal setae. Subanal scale suboval with 1+1 setae at caudal margin.

Gonopods: Anterior gonopods with T-shaped coxal part of coxosternum; telopodite flagelliform, positioned on posterior face of posterior gonopod colpocoxites inside a narrow deep curved sheath groove ( Fig. 3 ). Distal part of telopodite extending well beyond colpocoxite, tip pointed, unmodified. Colpocoxites broad, fused basally, each with a small excavation in curved posterad distal part and with lateral branch connected with angiocoxite frontally as well. Mesal sheath processes fused medially into a single lamina with medial outgrowth. Lateral sheath processes subtrapeziform. Angiocoxite with conical projection in posterior surface. Anterior angiocoxal process large, passing through foramina in colpocoxite ( Fig. 4 ). Distal part of anterior angiocoxal process with subapical small tooth ( Fig. 5 ). Posterior angiocoxal process absent. Telopodites of posterior gonopods 2-segmented, setose; femur short.

Female. Length 16–16.5 mm, width 1.8–1.9 mm with and 1.3–1.4 mm without paraterga. Body with 29 segments, ocelli 22–23. The spinicle at base of claws shorter than in male. Claws of legs 1 and 2 each with two dorsal and one ventral accessory claws. Other nonsexual characters as in male. Vulvae not dissected for examination.

Remarks: It is the fourth species of the genus Tokyosoma VERHOEFF, 1932 distributed in Japan and Korea (MIKHALJOVA 2000). The species Tokyosoma ronkayi ( SHEAR, 1990) (= Diplomaragna ronkayi SHEAR, 1990 ), formerly only known from North Korea, has recently also been recorded in South Korea ( MIKHALJOVA & LIM 2000). However, a study of the holotype of this species housed in the Hungarian Natural History Museum proved that it was a misidentification. The material from South Korea mentioned in the paper above appears to belong Tokyosoma hallum sp. n.

Notably, the re-examination of Tokyosoma ronkayi holotype has revealed the body consisting of 29 segments. The original description of Tokyosoma ronkayi erred in ascribing 32 body segments to that species (cf. SHEAR 1990).

In addition, the type material of Pterygostegia korsosi ( SHEAR, 1990) (= Diplomaragna korsosi SHEAR, 1990 ) housed in the HNHM was re-examined, too. The body of both male-holotype and females-paratypes is composed of 29 segments, not 32 as described in original description (cf. SHEAR 1990). Hence, the original diagnosis of these species must be corrected by the entry of 29 body segments instead of 32 somites.

*

Orientyla dahurica ( GERSTFELDT, 1859) – 1 m – North Korea, Prov. Ryanggang, Konchang , 800 m, No. 1369., 30.June 1988., leg. MERKL O. & SZÉL GY.

Remarks: This species, orginally described as Craspedosoma dahuricum from near the mouth of Shilka River, Chita Area, Siberia ( GERSTFELDT 1859), is a senior subjective synonym of Diplomaragna mikhaljovae SHEAR, 1990 , a species described from the southern part of the Russian Far East (Amurskaya Area, Primorsky kray = Maritime Prov.) ( SHEAR 1990), and later transferred to Orientyla (MIKHALJOVA 2000) . The synonymy has been established by MIKHALJOVA and GOLOVATCH (2001). This is the first record of O. dahurica in Korea.

Maritimosoma turova ( MIKHALJOVA, 1997) – 1 m, 1 f – Russia, Far East, Primorsky kray, Ussuriysky Nature Reserve, hills on the left side of river Komarovka , 200–500 m a.s.l., No. 39., 12.July 1990., leg. SZIRÁKI GY .

Remarks: This species, originally described in Diplomaragna ( MIKHALJOVA 1997) , and later transferred to Maritimosoma (MIKHALJOVA 2000) , is currently known only from Primorsky kray (= Maritime Prov.), its terra typica.

Figures

Figs 1–5. Tokyosoma hallum sp. n.: 1 = legpair 10, caudal view, 2 = legpair 11, frontal view, 3–4 = gonopod, caudal and frontal view, 5 = anterior angiocoxal process, frontal view. Scale in mm