Chlaenius (Lithochlaenius) noguchii Bates, 1873

Valim, Ye, Kavanaugh, David H., Shi, Hongliang & Liang, Hongbin, 2011, A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species, ZooKeys 128, pp. 15-52 : 30-31

Chlaenius (Lithochlaenius) noguchii Bates Figs 28-29 58 72 86 105-106 129-130 158

Chlaenius noguchii Bates, 1873:251; Chaudoir, 1876:192; Kryzhanovskij, 1976:12; Kirschenhofer, 1997:116; Kirschenhofer, 2005:491

Specimens examined.

Total 47 specimens: Japan: 10 males and 11 females (IZCAS), "Shiga, Takashima, Biwa Lake, 2008.3.30, Liang H.B. coll."; 1 male and 1 female (IZCAS), "Yadorigi-zawa, Tanzawa, Kanagawa"; 1 male (IZCAS), "Sado-shima Island"/ " 1968.VIII.20"; 2 males and 2 females (CASC), "Harima Japan May 1916"; 1 female (CASC), "Hozukyo Kyoto 1951.9.21"; 4 males and 2 females (CASC); "Kobe V-1912"/ "L. Gressitt Collector"; 1 female (CASC), "Kyoto, Yase, Oct. 21, 1951, Col. T. Horio"; 2 females (CASC), "Mimasaka Japan VII. J. E. Lewis"; 2 males and 2 females (CASC), "Mimasaka Japan VII-1912"/ "Coll. by J. C. Thompson"; 1 male and 1 female (CASC), " Nikko Japan VII.30.23"/ "ECVan Dyke collector"; 2 females (CASC), "Nikko Hondo Japan VII-1912"/ "Coll. by J. C. Thompson"; 1 female (CAS), Tokyo, Japan VI-6-31."/ "L. Gressitt Collector".

Diagnosis.

Antennomere 1 coniform ( Fig. 58 ); intervals 1-7 strongly convex, carinate, glabrous medially ( Fig. 86 ).

Description.

Total length = 14.50-16.00 mm (mean = 15.33), width = 5.07-5.60 mm (mean = 5.33); HW = 2.50-2.90 mm (mean = 2.67), EYL = 1.00-1.10 mm (mean = 1.02), ratio Ant3/Ant1 = 1.56-1.88 (mean = 1.72), PL/PW = 0.83-0.90 (mean = 0.86), EL/EW = 1.75-1.91 (mean = 1.80), EL/ PL = 1.29-1.40 (mean = 1.35).

Head and pronotum black with strong green or bluish green metallic luster; elytra black with very weak blue luster; ventral surface black; legs, antennae, mandibles and palpomere dark brown.

Head with vertex coarsely punctate behind eyes; antennomere 1 coniform ( Fig. 58 ); clypeus and labrum nearly truncate at apex; mentum tooth emarginate. Pronotum with disk finely punctate, nearly glabrous; basal foveae deep, finely pubescent and punctate (punctations much smaller than those on vertex). Elytra with intervals strongly convex, intervals 1-7 carinate, smooth and glabrous medially, with a row of punctation laterally ( Fig. 86 ), intervals 8-9 flat, densely pubescent and punctate; striae deep, punctate; humeral angle obtuse ( Fig. 72 ). Abdominal sterna IV-VI sparsely pubescent medially; apex of sternum VII more distinctly truncate in female than in male ( Figs 105-106 ). Aedeagus slender, flattened ( Fig. 130 ), lamella broadened to the left side, asymmetric ( Fig. 129 ), thin and bent ventrally ( Fig. 130 ).

Geographical distribution.

Fig. 158 . Confirmed only from Japan.

Remarks.

Mensural data cited in the description were based on measurements obtained from 5 males and 5 females selected for maximum variation.

This species may well be endemic to Japan, and its reported occurrence in Korea ( Kwon and Lee 1986) and Vietnam ( Paik et al. 2006) requires further clarification. The slender aedeagus of males of this species is very different from the stout aedeagi of other species of the agilis group.

Figures

Figure 158. Map showing the known geographical distributions of Chlaenius (Lithochlaenius) spp.

Figures 28 - 33. Chlaenius (Lithochlaenius) spp 28 Chlaenius noguchii Bates, male, in IZCAS, dorsal view 29 Chlaenius noguchii Bates, male, in IZCAS, ventral view 30 Chlaenius wrasei Kirschenhofer, male, in IZCAS, dorsal view 31 Chlaenius wrasei Kirschenhofer, male, in IZCAS, ventral view 32 Chlaenius agiloides Jedlicka, male, in IZCAS, dorsal view 33 Chlaenius agiloides Jedlicka, male, in IZCAS, ventral view. Scale line = 5.0 mm.

Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm.

Figures 66 - 79. Humeral region of elytra of Chlaenius (Lithochlaenius) spp 66 Chlaenius chuanqianensis sp. n., holotype 67 Chlaenius linwensini sp. n., holotype 68 Chlaenius propeagilis sp. n., holotype 69 Chlaenius agilis Chaudoir, holotype 70 Chlaenius formosensis Lorenz, CCCC 71 Chlaenius leishanensis Kirschenhofer, IZCAS 72 Chlaenius noguchii Bates, IZCAS 73 Chlaenius wrasei Kirschenhofer, IZCAS 74 Chlaenius agiloides Jedlicka, IZCAS 75 Chlaenius rambouseki Lutshnik from Ussuri region, ZCAS 76 Chlaenius anchomenoides Bates, paratype 77 Chlaenius nuristanus Jedlicka, paratype 78 Chlaenius nuristanus Jedlicka, paratype 79 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm.

Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm.

Figures 94 - 116. Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species 94 Chlaenius chuanqianensis sp. n., male, holotype 95 Chlaenius chuanqianensis sp. n., female, paratype 96 Chlaenius linwensini sp. n., male, holotype 97 Chlaenius linwensini sp. n., female, paratype 98 Chlaenius propeagilis sp. n., male, holotype 99 Chlaenius propeagilis sp. n., female, paratype 100 Chlaenius agilis Chaudoir, male, holotype 101 Chlaenius formosensis Lorenz, male 102 Chlaenius formosensis Lorenz, female 103 Chlaenius leishanensis Kirschenhofer, male 104 Chlaenius leishanensis Kirschenhofer, female 105 Chlaenius noguchii Bates, male 106 Chlaenius noguchii Bates, female 107 Chlaenius wrasei Kirschenhofer, male 108 Chlaenius wrasei Kirschenhofer, female 109 Chlaenius agiloides Jedlicka, male 110 Chlaenius agiloides Jedlicka, female 111 Chlaenius rambouseki Lutshnik, from Ussuri region, male 112 Chlaenius rambouseki Lutshnik from Ussuri region, female 113 Chlaenius anchomenoides Bates, male, paratype 114 Chlaenius nuristanus Jedlicka, male, paratype 115 Chlaenius nuristanus Jedlicka, female, paratype 116 Chlaenius formosanus Jedlicka from Taiwan, male. Scale line = 1.0 mm.

Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130).