Bembidion (Zemetallina) chalceipes Bates, 1878

Larochelle, André, Zeperyphodes, Broun & Larivière, Marie-Claude, 2015, Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini), Insecta Mundi 2015 (415), pp. 1-78 : 45-46

Bembidion (Zemetallina) chalceipes Bates, 1878

Fig. 35 , 68 , 78

Bembidium chalceipes Bates, 1878b: 24 . Type locality: West Coast, South Island.

Other synonymy as in Larochelle and Larivière (2001: 81–82).

Description. Body length 4.4–5.2 mm. Black; antennae black (segment 1 rufous underneath, segments 2–3 rarely rufous underneath); legs black to piceous (tibiae rarely rufous). Microsculpture very strong, isodiametric. Shiny, with strong metallic lustre (aeneous, rarely greenish); tibiae sometimes with slight metallic lustre. Forebody moderately wide in comparison to elytra. Head. Antennae filiform, moderately long (reaching about elytral shoulders). Thorax. Pronotum strongly convex, very wide; sides strongly rounded anteriorly, moderately sinuate posteriorly; posterolateral angles rectangular, vaguely obtuse at tip; laterobasal foveae moderately deep, linear, moderately long, reaching or almost reaching basal margin. Epipleura (in dorsal view) exposed in front of posterolateral angles. Elytra. Moderately convex (somewhat less than in B. anchonoderus ), subovate, moderately wide, widest about middle. Shoulders moderately rounded. Sides moderately rounded. Scutellar striole consisting of a row of punctures. Striae 2–3 complete, 4–6 incomplete; striae shallow, finely punctate; stria 7 obsolete, incomplete, at most with rudimentary punctures. Intervals depressed. Apical striole shallow, connected to stria 5; preapical setiferous puncture not isolated. Abdomen. Last visible sternum (sternum VII) of female with four long ambulatory setae only. Aedeagus. Lateral view ( Fig. 68 ): strongly arcuate, gradually narrowed from base to apex; base moderately concave dorsally; middle strongly convex dorsally, moderately concave ventrally; apex triangular, with extreme tip moderately wide and long.

Material examined. 301 specimens ( JNNZ, MONZ, NZAC).

Geographic distribution ( Fig. 78 ). South Island: BR, CO, DN, FD, KA, MB, MC, MK, NC, NN, OL, SC, SD, SL, WD.

Ecology. Lowland, montane, subalpine. Fossorial, epigean. Banks and beds of rivers and brooks, at a certain distance (1–4 m) from water. Open ground; moist or wet, sandy, bare or sparsely vegetated soil, sometimes mixed with silt. Nocturnal; hides during the day in the sand and under small stones. Gregarious.

Biology. Seasonality: September, November–April. Tenerals: February–March. Often infested with fungi (Laboulbeniales). Defence mechanism: when alarmed, the adult escapes by running.

Dispersal power. Macropterous, probably capable of flight. Fast runner. Vagility likely favoured by flight capacity.

Collecting techniques. Pouring water over the ground; treading the soil with the feet; turning small stones.

Reference. Larochelle and Larivière 2001: 81 (catalogue; biology, dispersal power, ecology, geographic distribution, references).

Figures

Figures 33–36. Dorsal habitus. 33) Bembidion bullerense new species. 34) B. hokitikense. 35) B. chalceipes. 36) B. wanakense. Scale line = 1 mm.

Figures 61–69. Aedeagi, lateral. 61) Bembidion parviceps. 62) B. waiho new species. 63) B. urewerense. 64) B. mangamuka new species. 65) B. solitarium. 66) B. bullerense new species. 67) B. hokitikense. 68) B. chalceipes. 69) B. wanakense.

Figures 77–82. Species distribution maps. 77) Bembidion callipeplum. 78) B. chalceipes. 79) B. chalmeri. 80) B. charile. 81) B. dehiscens. 82) B. embersoni.