Bembidion (Zemetallina) anchonoderus Bates, 1878

Larochelle, André, Zeperyphodes, Broun & Larivière, Marie-Claude, 2015, Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini), Insecta Mundi 2015 (415), pp. 1-78 : 37-38

Bembidion (Zemetallina) anchonoderus Bates, 1878

Fig. 26 , 59 , 74

Bembidium anchonoderum [sic] Bates, 1878a: 195. Type locality: Tairua, CL.

Other synonymy as in Larochelle and Larivière (2001: 81).

Description. Body length 4.0– 4.8 mm. Black; elytra piceous; antennae (segment 1 rufous, at least underneath) and legs piceous (tibiae rarely dark rufous). Microsculpture very strong, isodiametric. Shiny, with strong metallic lustre (aeneous, sometimes greenish or bluish); tibiae sometimes with slight metallic lustre. Forebody moderately wide in comparison to elytra. Head. Antennae filiform, moderately long (reaching about elytral shoulders). Thorax. Pronotum moderately convex, moderately wide; sides moderately rounded anteriorly, moderately sinuate posteriorly; posterolateral angles slightly projected laterally, somewhat obtuse at tip; laterobasal foveae moderately deep, linear, moderately long, not reaching basal margin. Epipleura (in dorsal view) exposed in front of posterolateral angles. Elytra. Moderately convex, subovate, moderately wide, widest about middle. Shoulders moderately rounded. Sides moderately rounded. Scutellar striole consisting of a row of punctures. Striae 2–6 complete, deep (even apically), coarsely punctate; stria 7 strong, incomplete, coarsely punctate. Intervals slightly convex. Apical striole deep, connected to stria 5 or 7; preapical setiferous puncture not isolated. Abdomen. Last visible sternum (sternum VII) of female with four long ambulatory setae only. Aedeagus. Lateral view ( Fig. 59 ): moderately arcuate, slender; base slightly convex dorsally; middle moderately convex dorsally, moderately concave ventrally; apex narrow, triangular, straight dorsally and ventrally, with extreme tip wide and long.

Material examined. 3,470 specimens ( CMNH, LUNZ, MONZ, NZAC).

Geographic distribution ( Fig. 74 ). North Island: AK, BP, CL, GB, HB, RI, TK, TO, WA, WI, WN, WO. South Island: BR, CO, DN, FD, KA, MB, MC, MK, NC, NN, OL, SC, SD, SL, WD.

Ecology. Lowland, montane. Epigean, fossorial. Banks and bars of rivers and brooks, at a certain distance (1–5 m) from water. Open ground; wet, fine sandy bare or sparsely vegetated soil, often with scattered stones and gravel. Nocturnal; hides during the day under stones, among gravel, in sand, at the base of plants, in soil crevices, in the soil, under fallen leaves and algal mats. Gregarious.

Biology. Seasonality: September–April, July. Tenerals: November–March. Occasionally infested with mites and fungi (Laboulbeniales). Defence mechanism: when alarmed, the adult escapes by running.

Dispersal power. Macropterous, probably capable of flight. Moderate runner. Vagility likely favoured by flight capacity.

Collecting techniques. Turning stones; raking the soil; pouring water over the ground; treading the soil with the feet.

Reference. Larochelle and Larivière 2001: 81 (as Bembidion (Zemetallina) anchonoderus ; catalogue; biology, dispersal power, ecology, geographic distribution, references).

Figures

Figures 25–28. Dorsal habitus. 25) Bembidion tekapoense. 26) B. anchonoderus. 27) B. stewartense. 28) B. parviceps. Scale line = 1 mm.

Figures 53–60. Aedeagi, lateral. 53) Bembidion musae. 54) B. orbiferum. 55) B. giachinoi. 56) B. callipeplum. 57) B. nesophilum. 58) B. tekapoense. 59) B. anchonoderus. 60) B. stewartense.

Figures 71–76. Species distribution maps. 71) Bembidion actuarium. 72) B. alacre. 73) B. albescens. 74) B. anchonoderus. 75) B. brullei. 76) B. bullerense new species.