Coprophanaeus (Coprophanaeus) conocephalus ( d'Olsoufieff , 1924)

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 66-68

Coprophanaeus (Coprophanaeus) conocephalus ( d'Olsoufieff, 1924) Plate 16A

Phanaeus (C.) conocephalus d’Olsoufieff, 1924: 72 (original description. Type locality: Equateur [= Ecuador], Loja).

Coprophanaeus conocephalus : Blackwelder 1944: 209 (list of species from Latin America); Arnaud 1982a: 116 (list of types at the MNHN).

Coprophanaeus (Coprophanaeus) conocephalus : Arnaud 2002a: 48 (diagnosis); Edmonds and Zídek 2010: 59 (characters in key), 69 (redescription); Carvajal et al. 2011: 320-321 (cited for Ecuador); Krajcik 2012: 204 (complete list of species, cited as subgenus of Phanaeus Macleay, 1819); Arnaud, 2018: 6 (comment), 7 (figure 2a); Chamorro et al. 2018: 93 (cited for Ecuador).

Phanaeus roubali Balthasar, 1939e: 241 (original description); Blackwelder 1944: 210 (list of species of Latin America); Arnaud 1997: 7 (synonym of Coprophanaeus conocephalus Ols); Arnaud 2002a: 48 (cited as synonym); Edmonds and Zídek 2010: 69 (cited as synonym); Bezdek and Hajek 2013: 433 (catalog of types NMPC).

Type specimens.

Phanaeus (Coprophanaeus) conocephalus d’Olsoufieff, 1924. The holotype (♀) is deposited at the MNHN. Locality: Equateur Loja, examined.

Holotype (♀): "Equateur / Loja / Abbé Gaujon [p, black margin]", "Ph. Conocephalus n. sp ♀ / det. G. Olsouffieff [hw and p]", "HOLOTYPE [p, red label]", "Coprophanaeus / conocephalus / Ols. / Holotype ♀ / P. Arnaud DET 1981 [p and hw]".

Phanaeus roubali Balthasar, 1939. The holotype (♂) is deposited at the NMPC (see Bezdek and Hajek 2013: 433) Locality: Ecuador, Lola [= Loja], not examined.

Distribution.

Only known from Ecuador.

Records examined.

BOLIVAR: Bosque Protector Filo Palanga, 970 m (1 specimen MUTPL). CAÑAR: Javin. La Trancel [= La Troncal], 850-1300 m (3 specimens CEMT; 1 specimen MQCAZ). GUAYAS: Cerecita Pta Chapella (3 specimens CEMT; 1 specimen MQCAZ). LOJA: without specific locality (1 specimen MNHN). MANABÍ: El Aromo, Río de los Napos, 280 m (1 specimen CEMT); El Aromo, 370 m (1 specimen MUTPL); Refugio de Vida Silvestre Pacoche, 340 m (3 specimens CEMT); Puerto López, Río Blanco, 272 m (1 specimen MUTPL).

Literature records.

GUAYAS: Balzar ( Edmonds and Zídek 2010: 71)

Temporal data.

Collected in January, February, April, August, and September.

Remarks.

Inhabits the coastal montane cloud forests and coastal evergreen foothill forests from 280-1300 m a.s.l. Collected with pitfall traps baited with carrion and human feces.