Bothynus

López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 456

Key to the species of adult Bothynus of Colombia

1 Head without central tubercle. Protibia tridentate ( Fig. 10 ). Elytra strongly punctate with deep punctures arranged in distinctive rows. Apex of metatibia truncate and without spinules. Length 14–20 mm. Parameres elongate, apices strongly dilated ( Figs. 12–13 )...................................................................... .. Bothynus herbivorus (Arrow)

- Head with central tubercle. Protibia quadridentate ( Fig. 18 ). Elytra nearly smooth, with very small, inconspicuous punctures. Metatibial apex with irregular margin and spinules. Length> 22 mm ............................................. 2

2 Males with protarsal inner claw simple (Fig. 7). Parameres elongate, apices strongly dilated, lobe-shaped (Figs. 4–5). Females can be only identified by association with males................................... Bothynus complanus (Burmeister)

- Males with protarsal inner claw with median tooth ( Fig. 23 ). Parameres shorter and wider, apices slightly dilated ( Figs. 20–21 ). Females can be only identified by association with males.......................... Bothynus quadridens (Taschenberg)

Figures

FIGURES 9 – 16. Bothynus herbivorus. 9) female, habitus in dorsal view, 10) female, habitus in frontal view, 11) head of male, 12) parameres in frontal view, 13) parameres in lateral view, 14) spiculum gastrale, 15) pygidium of female, 16) locality records in Colombia.

FIGURES 17 – 24. Bothynus quadridens. 17) habitus in dorsal view, 18) habitus in frontal view, 19) head of male, 20) parameres in frontal view, 21) parameres in lateral view, 22) spiculum gastrale, 23) male inner proclaw, 24) locality records in Colombia.