Goja risaraldensis, Carrejo & Obando & Casasola-González & García Aldrete, 2021

Carrejo, Nancy, Obando, Ranulfo González, Casasola-González, José Arturo & García Aldrete, Alfonso N., 2021, New Colombian Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico, Zootaxa 5040 (4), pp. 451-481 : 465-468

Goja risaraldensis n. sp. Male

( Figs 19–24 )

Diagnosis. Similar to P. andina n. sp., from which it differs in the wing venation, in having larger external parameres, in the head pattern, and in the endophallic sclerites (Compare Figs 1–2 with 19–20, 3 with 21, and 6 with 24 in this paper). Apex of the aedeagal arch robust, with the posterior border concave and a pair of the left endophallic sclerites more widened distally than in P. andina ( Fig. 24 ).

Color (in 80% ethanol). Body brown with light spots. Head pattern ( Fig. 21 ); forehead brown, with broad ochre band between the compound eyes, enclosing the ocellar group. Compound eyes dark brown; ocelli hyaline, with ochre centripetal crescents. Anteclypeus brown, with ochre striae converging mesally as illustrated; labrum dark brown; genae cream to light brown, with ochre anterior areas; pre- and postgenitalia cream. Base of maxillae cream, with brown spots, maxillary palps brown, Mx4 darker.Antennae light brown. Thorax light brown, pronotum cream, tergal lobes of meso- and metathorax brown; pleura cream, with ochre spots. Legs light brown, coxae with basal ochre spots. Wings hyaline, veins light brown. Forewings ( Fig. 19 ) with abundant brown setae. Abdomen cream, with ochre brown subcuticular spots. Clunium, epiproct and paraprocts light brown, with brown spots. Hypandrium light brown. Phallosome proximally creamy, distally brown; aedeagal arch and endophallic sclerites dark brown, external parameres brown.

Morphology. Head: H/MxW: 1.48; compound eyes small, H/d: 3.75; IO/MxW: 0.61. Outer cusp of lacinial tips broad, with seven denticles. Mx4/Mx2: 1.15. Labrum elongate, lower margin with a pair of projections with rounded apex, with five distal sensillae, one central placoid, flanked by a pair of trichoid-placoid. Forewings ( Fig. 19 ): FW/W: 2.81; pterostigma elongate, wider in the middle: lp/wp: 6.04; Rs with seven branches; areola postica elongate: al/ah: 4.32; M with six branches, M5 and M6 forked. Hindwings ( Fig. 20 ): HW/w: 3.12. Rs and M with four branches. Legs: Fore- tarsi without ctenidiobothria, mid- and hind- tarsi with ctenidiobothria. Hypandrium ( Fig. 23 ) broad, setose, posterior border slightly concave in the middle. Phallosome ( Fig. 24 ): deeply obtusely concave anteriorly, side struts slender, straight; aedeagal arch robust, apically with two short projections, directed outward; external parameres well developed, inner border almost straight, outer border rounded, right one distally acuminate, left one blunt distally. Three pairs of asymmetric endophallic sclerites, as illustrated ( Fig. 24 ). Clunium projecting posteriorly over the area of the epiproct. Paraprocts large, broadly elliptic ( Fig. 22 ), setae on posterior border as illustrated with setae and macrosetae; large elliptic sensory fields with 32-33 trichobothria in basal rosettes. Epiproct convex anteriorly, sides converging to almost straight posterior border; four macrosetae mesally, other setae as illustrated ( Fig. 22 ) broad.

Measurements. FW: 4500, HW: 3275, F: 1080, T: 1980, t1: 995, t2: 280, ctt1: 39, f1: 1020, f2: 960, f3: 780, Mx4: 390, IO: 295, D: 252, d: 315, IO/d: 1.55, PO: 0.85.

Specimen studied. Holotype male. COLOMBIA. Risaralda, Santuario, Planes de San Rafael , 5º07’32.4”N: 76º00’32.4”W, 2099 m. 18.VII.2015, on tree trunk. R. González, N. Calderón and O. Saenz.

Etymology. The specific epithet refers to the department of Risaralda, Colombia, where the holotype was found.

Figures

FIGURES 19‒24. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm.

FIGURES 1‒6. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm.