Collita vietnamica, Dubatolov, 2012

Volynkin, Anton V., Černý, Karel, Huang, Si-Yao & Hu, Hua-Lin, 2023, Taxonomic review of the Oriental genus Gandhara Moore with descriptions of four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini), Zootaxa 5374 (3), pp. 390-408 : 396

The G. vietnamica species-group

Diagnosis. The male genital capsule of the species-group is characterised by the elongate and narrow valva with almost parallel margins medially and being tapered only in its distal third. Additionally, compared to the G. typica species-group, the sacculus of the G. vietnamica species-group is equal in length to the dorsal section of the valva (it is shorter than the dorsal section in the G. typica species-group), and the uncus is uniformly slender whereas it is thicker and distally dilated in the G. typica species-group. The lobes of the anellus of the species-group are elongate and plank-like whereas they are shorter and triangular in the G. typica species-group. The phallus of the G. vietnamica species-group is markedly shorter than in the G. typica species-group, and its vesica is shorter, lacking the proximal tubular section but having larger utricular diverticula, the distal of which bears a conical terminal cornutus, which is claw-shaped in the G. typica species-group. In the female genitalia, the species-group differs from the G. typica species-group in the horseshoe-shaped antrum (it is funnel-shaped in the G. typica species-group), the markedly shorter posterior section of the corpus bursae having heavy sclerotisation only on the left side and bearing an appendix bursae directed laterally on the right side (it is directed postero-laterally in the G. typica species-group). Additionally, the anterior section of the corpus bursae of the G. vietnamica species-group is broader and bears a signum medially whereas it is situated more posteriorly in the G. typica species-group.