Holzenthal, Ralph W., Blahnik, Roger J. & Rios-Touma, Blanca, 2018, New species and a new genus of Philopotamidae from the Andes of Bolivia and Ecuador (Insecta, Trichoptera), ZooKeys 780, pp. 89-108: 89

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gen. n.

Aymaradella   gen. n. Figs 1, 2, 3

Type species.

Aymaradella boliviana   , new species


Aymaradella   , new genus, can be distinguished from any other genus of Philopotaminae  by the loss of 2A vein in the hind wing, the synscleritous tergum and sternum of segment VIII, and the elongate sclerotized dorsal processes of segment VIII.

This species has the general venational attribute of lacking the second anal vein in the hind wing, a character used to define the genera Wormaldia   and Gunungiella   . However, it is very distinctly different from either of those genera in overall form, and completely unlike any described species of Wormaldia   from either North or South America. In particular, the completely fused segment VIII, with elongate dorsal processes, is unique among species in Philopotaminae  .


Adult. Head relatively short and rounded, postparietal sclerite short (ca. ½ diameter of eye). Spur formula 2:4:4. Maxillary palps 5-segmented, segment II with apicomesal bristles, labial palps 3-segmented. Venation complete for Philopotamidae   (forewings with forks I-V, hind wing lacking fork IV); forewing with discoidal cell relatively short, forks I and II approximately sessile, crossveins s, r-m, and m hyaline and nearly linear, 3A looped to 2A, 2A to 1A, intersecting in proximal half of vein. Fork I of hind wing with short stem, fork II sessile, 1A and 3A intersecting wing margin, 2A missing.

Male. Sternum VII with short, rounded mesoventral process (rather than flattened, spatulate process typical of Wormaldia   ). Segment VIII synscleritous, expanded anterodorsally, dorsal margin with pair of elongate processes. Segment IX synscleritous, narrowing dorsally. Tergum X simple in structure and entire, wide basally, narrowing apicaly, with numerous sensilla and/or short setae. Preanal appendages elongate, narrow, digitiform, emerging near base of tergum X. Inferior appendages bi-segmented, linear, apical segment with mesal pad of short stiff setae. Phallic apparatus simple in structure, without spines or inclusions.

Female. Genitalia elongate and tapering from segment VII; segment VII longer than preceding segment, with small ventral process at midlength; segment VIII nearly as long as segment VII, tapering, not synscleritous, sternite with lateral pair of very elongate, narrow apodemes. Segment IX very short, anterolateral margin with pair of elongate, narrow apodemes, extending anterad. Segment X composed of pair of elongate, narrow sensillate lobes, each with short apical cercus. Vaginal apparatus membranous, without evident sclerites.


The name Aymara is considered feminine and refers to the people and language of Bolivia; the suffix is considered to be a diminutive and makes the name euphonious with Chimarrhodella   and Hydrobiosella   , other philopotamids previously and newly known from the Neotropics.