Jungiella syriaca Omelková & Ježek,

Omelková, Markéta & Ježek, Jan, 2017, Two new species of Jungiella (Diptera: Psychodidae: Psychodinae) from the Palaearctic Region, Zootaxa 4250 (6), pp. 560-576: 569-574

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Jungiella syriaca Omelková & Ježek

sp. nov.

Jungiella syriaca Omelková & Ježek  sp. nov. ( Figs 21–42View FIGURES 21 – 26View FIGURES 27 – 34View FIGURES 35 – 37View FIGURES 38 – 42)

Type locality. Syria, Prov. Al Lathiqiyeh: a streamlet NW Rabi'ah, 35°49'N, 36°02'E.GoogleMaps 

Type material. Holotype: male, dissected on slide, Syria, Prov. Al Lathiqiyeh: a streamlet NW Rabi'ah, shaded by swampy vegetation, forested and shrubbery region with many hills, 35°49'N, 36°02'E, SW; 1.– 2.vii.1998, P. Chvojka leg; Cat. No. 34656, Inv. No. 21967.GoogleMaps 

Paratypes: 3 males, dissected on slides, data same as in holotype, Cat. No. 34657–34659, Inv. No. 21968– 21970.

Description. Male. Head ( Fig. 21View FIGURES 21 – 26) 0.9 times as long as broad, vertex little swollen on either side, sides subtending obtuse angle, no median scar-free band, occipital lobe elevated, prominent. Corniculae inflated distally ( Figs 21, 22View FIGURES 21 – 26), club-shaped, 1.4 times as long as distance between both bases of first palpomeres. Eye bridge of 4 facet rows ( Figs 21View FIGURES 21 – 26, 27View FIGURES 27 – 34), separated by distance equal to 1.5–1.9 facet diameters, near frontal suture little more. The ratio of distance of apices of eyes (tangential points) to minimum width of frons approximately 7.7:1. Interocular frontal suture ( Figs 21View FIGURES 21 – 26, 27View FIGURES 27 – 34), with thin convex ligament, conspicuously sclerotized, U-shaped, median rib inverted to vertex. Frontoclypeus ( Fig. 21View FIGURES 21 – 26) with large basal oblong scar patch merging into thin irregular median band extending nearly to frontal suture. Insertions of postocular bristles on dorsal apices of eyes enlarged ( Fig. 21View FIGURES 21 – 26).

Antennae ( Figs 23View FIGURES 21 – 26, 28View FIGURES 27 – 34) incomplete, with numerous microtrichiae, scape prolonged, cylindrical, little widened distally (1.4 times broader than the basis), 2.1 times longer than spherical pedicel. Flagellomeres amphora-shaped, asymmetrical, necks shorter than swollen basal parts. Sensory filaments (ascoids) imitating thin crescent-shaped rolls ( Figs 23View FIGURES 21 – 26, 28View FIGURES 27 – 34), as long as or little longer than flagellomeres, paired. Relative length of palpomeres 1.0:1.3:1.4:2.1 ( Fig. 29View FIGURES 27 – 34), last segment annulated. Maxilla 1.2 times shorter than first segment, mouthparts extend beyond ends of basal segments of maxillary palps ( Figs 21View FIGURES 21 – 26, 30View FIGURES 27 – 34). Terminal lobes of labium ( Fig. 24View FIGURES 21 – 26) without lines of spines between both folds. Relative ratio of maximum length of cibarium to length of epipharynx 1.9:1 ( Fig. 31View FIGURES 27 – 34).

Thorax. Shape of thoracic sclerites and insertions of macrosetae as in Fig. 32View FIGURES 27 – 34. Length ratios of femora, tibiae and first tarsomeres: P1 1.6:2.2:1.0, P2 1.8:3.1:1.3 and P3 2.0:2.8:1.3, paired tarsal claws of P1 elongated and bent subapically, haired in their basal part ( Fig. 25View FIGURES 21 – 26).

Wings ( Fig. 35View FIGURES 35 – 37) elongate, translucent, 1.7 mm long (paratypes 1.5–1.6 mm), with tendency to expand in anal and humeral regions, apically rounded, 2.8 times as long as its wide. Sc longer than basal cell, straight, strengthened continuously, tapering apically. R1 without streak, in contrast to J. janiki  . Next strengthened veins: R1 sometimes distally, R2, R4 only in basal cell, R5, M1+ 2 in basal cell, CuA1 and CuA2 (least conspicuously expanded in origin). Radial fork complete, medial fork incomplete and end of CuA2 incomplete in the holotype, paratypes have all three mentioned points complete. All type material has wing forks and end of CuA2 approximately at same level. R5 ends beyond wing apex. Bases of M3, CuA1 and CuA2 not connected. Halteres ( Fig. 33View FIGURES 27 – 34) stick-shaped, knobs covered with pedunculate scales; ratio maximum length to maximum halter width 2.9:1.

Male genitalia with ejaculatory apodeme prolonged, almost rectangular, with tongue-shaped ending proximally from dorsal view and stick-shaped from lateral view ( Figs 26View FIGURES 21 – 26, 36View FIGURES 35 – 37, 38View FIGURES 38 – 42). Aedeagal complex  dorsoventrally flattened, spatulate structures (parameral sheath) with paired sclerotized lamellae (distiphalus) jointed with two rounded „glenoid blades“ with glenoid facet on inner side and characteristic crescent cavities distally ( Figs 36View FIGURES 35 – 37, 38, 39View FIGURES 38 – 42) divided by shallow cleft between them. Furca wide U-shaped ( Fig. 34View FIGURES 27 – 34). Gonocoxites ( Figs 38, 40, 41View FIGURES 38 – 42) cylindrical, broader basally. Gonostyli thin, somewhat bent, gradually tapering to the end, approximately 1.5 times as long as gonocoxites. Epandrium ( Figs 37View FIGURES 35 – 37, 42View FIGURES 38 – 42) bare proximally, setose on both sides of shallow concave epandrial notch. Middle aperture ovoid, considerably sclerotized. Remainder of ventral epandrial plate well bordered. Hypandrium ( Figs 37View FIGURES 35 – 37, 42View FIGURES 38 – 42) forming semicircular curve with conspicuous widening in the middle. Epiproct largely tongue-shaped, hypoproct 1.8 times narrower, both parts of same length, setose. Surstyli ( Figs 37View FIGURES 35 – 37, 42View FIGURES 38 – 42) 1.5 times as long as epandrium, slightly C-shaped, subapically with stable number of 10 retinaculi gradually shorter towards top of surstylus. Retinacula not apically frayed ( Fig. 37View FIGURES 35 – 37).

Female unknown.

Differential diagnosis. Jungiella syriaca  sp. nov. differs from J. procera Krek, 1971  by the 2.1:1.0 length ratio of scape and pedicel ( Fig. 23View FIGURES 21 – 26), the position of medial wing fork (complete or incomplete), which is approximately at the line formed by the radial fork and end of CuA2 (complete or incomplete) ( Fig. 35View FIGURES 35 – 37). Further, Jungiella syriaca  sp. nov. has hypandrium with a conspicuous widening in the middle ( Figs 36View FIGURES 35 – 37, 38View FIGURES 38 – 42); the ejaculatory apodeme has ovoid aperture distally ( Figs 36View FIGURES 35 – 37, 38View FIGURES 38 – 42) and rounded „glenoid blades“ of male genitalia have crescent cavities distally ( Figs 36View FIGURES 35 – 37, 38, 39View FIGURES 38 – 42). J. procera  has length ratio of scape and pedicel 3.1:1.0, medial fork situated basal to the line formed by radial fork and end of CuA2 (medial fork and ending of CuA2 are complete), hypandrium without a protuberance in the middle, ejaculatory apodeme without aperture distally and rounded blades of male genitalia with tridentate sclerites distally. Both species belong to the subgenus Psychocha Ježek, 1983  . Etymology. The species was named after the country where the type specimens were collected. Bionomics. Unknown.

Distribution. Known only from the type serie collected in Syria.