Chimarra kadavuensis, Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58 : 43-44

Chimarra kadavuensis , new species

Figs. 24 , 129–133

Chimarra kadavuensis is unique among Fijian Chimarra species in the structure of the male genitalia, particularly the very short segment IX, long and narrow gonopods, and in the length of the phallic apparatus, which is shorter than the combined length of segment IX and the gonopods.

Male. Body brown, dorsal part of meso- and metathorax dark brown. Large dark area present between lateral and anterior ocelli. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 24 ). Forewings 6.4 mm (n=1), brown, iridescent; relatively broad, ratio of length to breadth 3.3; R1 undulating before crossvein r; radial sector not produced anterad immediately before discoidal cell; discoidal cell originating slightly distally of mid-length of wing, 2x longer than wide; median cell slightly longer than discoidal cell; crossvein r situated at base of R2; fork I originating tangentially with crossvein s; nygma located near base of fork II; fork III 1 /7th as long as wing; fork V as long as fork I, shorter than fork II; Cu2 well separate from A at wing margin. Hind wings 5.1 mm (n=1), brown, iridescent; broad, ratio of length to breadth 2.8; margin slightly incurved where Cu1 and Cu2 fused with margin; fork III as long as discoidal cell and 1/9 as long as wing; fork I originating from anterodistal corner of discoidal cell, about as long as fork V; 1A+2A 3x longer than 1A.

Male genitalia ( Figs. 129–133 ). Segment IX about 1.5x taller than long; ventral and dorsal margin about equally long in lateral view; segment narrowest immediately immediately below cerci; ventral anterior plates rounded anteriorly; dorsal anterior plates short, slender, anteriorly rounded; anterodorsal margin widely and deeply concave in lateral view; ventral margin nearly straight; posterior margins weakly undulating, nearly vertical, curved anterad immediately below cercus. In dorsal view segment IX with short, narrow anterior lobes; anterodorsal margin forming deep, straight-margined incision, with anterad-orienting processes on posteromesal margins. In ventral view segment IX with undulating lateral margins; anterior and posterior margins concave; posterior margin without central projection. Tergum X divided into lateral branches at basis. Each lateral branch of tergum X with triangular, nearly straight dorsal branch pointing posterad in lateral view, with dorsal margin on line with dorsal margin of segment IX; ventral branch originating well below dorsal branch, slender, curved posterad along its length, with apical and subapical sensilla, reaching as far posterior as dorsal branch; small rounded plate present at base between dorsal and ventral branches. In dorsal view dorsal branches of tergum X narrowing distally, curved laterad along their length; ventral branches nearly parallel, each with laterad-curved apex. Cerci small, located near dorsal margin of segment IX and tergum X; directed posterad in lateral view, tube-shaped; covered by long setae. Gonopods narrow, tapering from basis to 3/ 4 in lateral view, curved posteroventrad along their length, about as long as height of segment IX. Gonopodsunbranched in lateral view; in ventral view narrow and curved mesad along their length; with well sclerotized, long mesal branches oriented posterad, nearly touching apically; megasetae absent. Phallic apparatus slightly shorter than rest of genitalia, straight, compact; phallotheca, in lateral view, with anterior part 3x thicker than narrowest posterior part; in ventral view anterior part nearly 3x wider than posterior part; apicoventral spine absent on phallotheca; phallotremal sclerite simple, 3-rayed; 2 nearly black endothecal spines present. Group of spicules absent.

Female. Unknown.

Holotype male: KADAVU: 0.25 km SW Solodamu Vlg., Moanakaka Bird Sanctuary, Malaise trap 4, 7.iii–11.iv.2004, 19.078°S, 178.121°E, 50 m, leg. E. Schlinger & M. Tokota’a [loc#46] [ FNIC].

Paratypes: Same data as holotype [loc#46] — 21 males [ NHRS]. Same data as holotype, except 19.xii.2003 – 18.i.2004 [loc#46] — 2 males [ BPBM]. Same data as holotype, except 11.iv–2.v.2004 [loc#46] — 28 males [ BPBM]. Same data as holotype, except 9–15.ii.2004 [loc#46] — 4 males [ NHRS]. Same data as holotype, except 15.ii–7.iii.2004 [loc#46] — 41 males [ NHRS, DNA voucher IN3]. Solodamu, Malaise trap in coastal limestone forest, 11.vi–5.vii.2003, 19°04’S, 178°07’E [19.0667°S. 178.1167°E], 128 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#47] — 3 males [ BPBM]. Same data, except 25.viii–23.x.2003 [loc#47] — 4 males [ NHRS]. Same data, except 23.x–19.xii.2003 [loc#47] — 2 males [ FNIC].

Etymology: Kadavuensis , named after the island Kadavu, the type locality of the species.

Distribution: Kadavu.

Figures

FIGURES 22 – 29. Right wings, dorsal views. 22 — C. cartwrighti, new species, holotype; 23 — C. kimminsi, new species, holotype; 24 — C. kadavuensis, new species, holotype; 25 — C. lavensis, new species, holotype; 26 — C. devoensis, new species, holotype; 27 — C. helomyzida, new species, holotype; 28 — C. tipulida, new species, holotype; 29 — C. psychodida, new species, holotype.

FIGURES 129 – 133. Chimarra kadavuensis, new species, holotype. 129 — genitalia, left lateral view; 130 — genitalia, dorsal view; 131 — genitalia, ventral view; 132 — phallic apparatus, left lateral view; 133 — phallic apparatus, ventral view.