Chimarra lavensis, Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58 : 44-46

Chimarra lavensis , new species

Figs. 25 , 134–138

The male genitalia of this species, particularly tergum X, resembles those of C. devoensis and C. vanuensis in the presence of a strong dorsad-oriented dorsal branch of tergum X. In C. lavensis the ventral branch of tergum X is shorter than the dorsal branch, while in the other 2 species the ventral branch is longer than the dorsal branch. The phallic apparatus of C. lavensis has no spicules, which are present in C. devoensis ; and the phallic apparatus of C. lavensis has 4 endothecal spines, whereas only 2 are present in C. vanuensis . The species is easily distinguished from C. vanuensis by the absence of a large pale spot between Cu1 and the end of Cu2 (arculus) in the forewings.

Male. Head and thorax brown, dorsal part of meso- and metathorax dark brown. Area between ocelli nearly black. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 25 ). Forewings 4.2 mm (n=1), brown. Forewings broad, ratio of length to breadth 3.1; R1 nearly straight before crossvein r; radial sector slightly produced anterad immediately before discoidal cell; discoidal cell originating at mid-length of wing, about 3x longer than wide; median cell slightly shorter than discoidal cell; crossvein r originating from basis of fork I; fork I originating before crossvein s at distance nearly equal to length of crossvein s; nygma located near base of fork II; fork III 1 /7th as long as wing; fork V slightly shorter than fork II; Cu2 ending in wing margin well separated from A. Hind wings 3.4 mm (n=1), brown; ratio of length to breadth 2.9; margin weakly incurved at arculus, where Cu1 and Cu2 fused with margin; fork I originating slightly before anterodistal corner of discoidal cell; fork III as long as discoidal cell and 1/11th as long as wing; fork V as long as fork I; 1A+2A about 3x as long as 1A.

Male genitalia ( Figs. 134–138 ). Segment IX nearly 2x taller than long in lateral view; anterodorsad produced into large, rounded, anterodorsad oriented plate; anteroventral plate large, triangular, slightly pointing dorsad in lateral view; anterodorsal margin widely and deeply concave; ventral margin irregularly convex, slightly incised at vertical apodeme; ventrum strongly produced anterad into triangular plate in lateral view; ventral margins irregularly convex; each posterior margin nearly straight; segment IX ventrally with with setae restricted to row before posterior margin ( Fig. 136 ). Dorsal part of segment IX slightly concave in lateral view, incised at vertical apodeme; anterior margin invisible in dorsal view. In ventral view segment IX with almost parallel lateral margins, slightly narrower at transverse apodeme; anterior margin nearly straight; posterior margin weakly concave, without central projection. Tergum X short, divided into pair of lateral branches; in lateral view each lateral branch with large dorsal, darkly pigmented, anteriorly hooked dorsal branch, and short tube-shaped, nearly straight ventral branch with 2 apical sensillae. In dorsal and ventral view, ventral processes oriented posterad, nearly parallel. Cerci minute, located at posterior margin of segment IX immediately above mid-height of tergum X; wart-like in lateral and dorsal view; covered by long setae. Gonopods longer than length of segment IX; divided into broad basal third and narrow distal 2/3rds in lateral view; distal 2/3rds nearly parallel-sided; each gonopod unbranched in lateral view; with well-developed dorsomesal lobe and ventromesal lobe of mesal process in ventral view. Anterodorsal margin of each gonopod weakly concave, smooth; posteroventral margin of basal 1/2 with strongly expanded setal bases, distal 1/2 with short setal bases; ventral margin convex; apex without megasetae. In ventral view, gonopods as narrow at base as rest of gonopods, with undulating margins; gonopods slightly curved mesad along their length. Mesal branch of each gonopod divided into well-developed ventromesal and dorsomesal lobe; ventromesal lobe strongly convex, darkly pigmented, covered by minute setae; dorsomesal lobe slightly curved ventrad, with rounded apex. Phallic apparatus about as long as rest of genitalia, straight along its length; phallotheca, in lateral view nearly 4x thicker than posterior part; in ventral view about 2x thicker than posterior part; apicoventral spine absent; phallotremal sclerite complex in lateral view; in ventral view forming single posterad directed arrow head; 4 nearly black, short, variously directed endothecal spines present, about as long as diameter of narrowest part of phallotheca; endothecal spicules absent.

Female. Unknown.

Holotype male: TAVEUNI: Cakaudrove Prov., Lavena, Malaise trap, 4.i–11.iii.2004, 16°51.315’S,

179°53.323’W [16.8553°S, 179.8888°W], 213 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#40] [ FNIC]. Etymology: Lavensis , after Lava Village, the type locality of the species. Distribution: Taveuni.

Figures

FIGURES 22 – 29. Right wings, dorsal views. 22 — C. cartwrighti, new species, holotype; 23 — C. kimminsi, new species, holotype; 24 — C. kadavuensis, new species, holotype; 25 — C. lavensis, new species, holotype; 26 — C. devoensis, new species, holotype; 27 — C. helomyzida, new species, holotype; 28 — C. tipulida, new species, holotype; 29 — C. psychodida, new species, holotype.

FIGURES 134 – 138. Chimarra lavensis, new species, holotype. 134 — genitalia, left lateral view; 135 — genitalia, dorsal view; 136 — genitalia, ventral view; 137 — phallic apparatus, left lateral view; 138 — phallic apparatus, ventral view.