Chimarra cartwrighti, Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58 : 40-41

Chimarra cartwrighti , new species

Figs. 22 , 117–123

The forewings of this species are dark brown and iridescent like those of C. manni . Chimarra cartwrighti is easily separated from C. manni by having male genitalia with segment IX appearing more or less rectangular in lateral view, and gonopods that are broader in lateral view.

Male. Head and thorax dark brown, dorsal part of meso- and metathorax even darker brown. Area between ocelli nearly black. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 22 ). Forewings 6.9 mm (n=1), dark brown, iridescent. Forewings broad, ratio of length to breadth 3.1; R1 weakly weaving before crossvein r; radial sector weakly produced anterad immediately before discoidal cell; discoidal cell originating distally of mid-length of wing, about 2x longer than wide; median cell slightly longer than discoidal cell; crossvein r originating from basis of R2; fork I originating before crossvein s at distance equal to 1/2 length of crossvein s; nygma located near base of fork II; fork III 1 /6th as long as wing; fork V longer than fork II; Cu2 ending in wing margin well separated from A. Hind wings 5.5 mm (n=1), brown, iridescent; ratio of length to breadth 2.8; margin weakly incurved at arculus, where Cu1 and Cu2 fused with margin; fork I originating from anterodistal corner of discoidal cell; fork III slightly longer than discoidal cell and 1/7th as long as wing; fork V longer than fork I; 1A+2A about 4x as long as 1A.

Male genitalia ( Figs. 117–123 ). Segment IX longer than high, nearly rectangular in lateral view; anterodorsal margins slightly produced into rounded lobe, ventrum slightly produced anterad and posterad in lateral view; ventral margins slightly convex; dorsal margins slightly concave; each posterior margin shallowly convex; segment IX ventrally with posterior third with setae ( Fig. 120 ). Dorsal part of segment IX membranous, expanded into 2 rounded, posterior lobes; anterior margin invisible in dorsal view. In ventral view segment IX with parallel lateral, undulating margins, anterior and posterior margin shallowly concave; posterior margin without central projection. Tergum X short, divided into pair of lateral branches; each branch with dorsal rounded lobe, and ventral, nearly straight, tube-shaped, posterad oriented ventral lobe with 2 apical sensillae. Tergum originating deeply inside segment IX. In dorsal and ventral view, tergum X forming pair of very slender processes separated by deep U-shaped gap; each process with slender, laterad oriented thorn-like outgrowth immediately basally of mid-length of each lateral branch. Cerci located at posterodorsal corner of segment IX; slender, slightly dilated and directed posterad in lateral view; broad, rounded in dorsal view; covered by long setae. Gonopods triangular in lateral view, narrowing uniformly apically along their length; about as long as height of segment IX;unbranched in lateral view; with well-developed dorsomesal lobe and ventromesal lobe of mesal process in ventral view. Dorsal margin nearly straight, smooth; posterior margin slightly concave, strongly undulating by large setal bases; ventral margin strongly convex; gonopods slightly longer than tergum X; apical margin armed with small megasetae. In ventral view, gonopods broad at base, slightly curved mesad toward apex; mesal margins strongly diverging in ventral view. Mesal branch divided into well-developed ventromesal and dorsomesal lobe; ventromesal lobe rounded convex, darkly pigmented, covered by minute setae; dorsomesal lobe strongly curved ventrad, with rounded apex. Phallic apparatus about as long as rest of genitalia, straight along its length; phallotheca, in lateral and ventral view about 2x thicker than posterior part; apicoventral spine absent; phallotremal sclerite minute, divided into 3 posterior rays; 2 nearly black, short, posterad directed endothecal spines present, slightly shorter than diameter of narrowest part of phallotheca; group of ventrad oriented spicules present distally in endotheca.

Female. Unknown.

Holotype male: VITI LEVU: Vuda Prov., Koroyanitu Pk., 1 km E Abaca Vlg., Malaise trap 1, 19–26.xi.2002, 17.667°S, 177.55°E, 800 m, leg. E. Schlinger & M. Tokota’a [loc#03] [ FNIC].

Paratypes: Same data as holotype, except 22.iv–5.v.2003 [loc#03] — 7 males [ NHRS]. Vuda Prov., Koroyanitu Pk., 1 km E Abaca Vlg., Kokabula Trail, Malaise trap, 12–19.xi.2002, 17°40’S, 177°33’E [17°40’S, 177°33’E 17.3333°S, 177.5500°E], 800 m, leg. E. Schlinger & M. Tokota’a [loc#05] — 1 male [ NHRS, DNA voucher IM8]. Naitasiri Prov., 4 km WSW Colo-i-Suva Village, Mt. Nakobalevu, Malaise trap 1, 12.iv.2004, 18.057°S, 178.42°E, 300 m, leg E. Schlinger & M. Tokota’a [loc#12] — 1 male [ BPBM]. Same data, except 9–30.v.2003, 18.055°S, 178.424°E, 372 m [loc#11] — 1 male [ NHRS].

Etymology: Cartwrighti , named after David I. Cartwright, for his valuable contribution in the taxonomy of Australian Chimarra .

Distribution: Viti Levu.

Figures

FIGURES 22 – 29. Right wings, dorsal views. 22 — C. cartwrighti, new species, holotype; 23 — C. kimminsi, new species, holotype; 24 — C. kadavuensis, new species, holotype; 25 — C. lavensis, new species, holotype; 26 — C. devoensis, new species, holotype; 27 — C. helomyzida, new species, holotype; 28 — C. tipulida, new species, holotype; 29 — C. psychodida, new species, holotype.

FIGURES 117 – 123. Chimarra cartwrighti, new species, holotype. 117 — genitalia, left lateral view; 118 — genitalia, dorsal view; 119 — tergum X, dorsal view; 120 — genitalia, ventral view; 121 — gonopods, caudal view; 122 — phallic apparatus, left lateral view; 123 — phallic apparatus, ventral view.