Chimarra psychodida, Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58 : 50-52

Chimarra psychodida , new species

Figs. 29 , 155–159

Chimarra psychodida resembles C. tipulida , C. malickyi , and C. levuensis in having gonopods each with both a slender, strongly produced dorsal branch, and a short, well-developed plate-like ventral branch. It is distinguished from C. malickyi in that segment IX is more rounded anteriorly in lateral view; the lateral branches of tergum X are nearly parallel; and tergum X is clearly shorter than each gonopod. It is separated from C. levuensis by the presence of parallel lateral branches of tergum X. It is separated from C. tipulida by the broader anterodorsolateral processes of segment IX; and slightly shorter and broader tergum X in lateral view.

Male. Body brown, dorsal part of meso- and metathorax dark brown. Large dark area between lateral and anterior ocelli. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 29 ). Forewings 5.7 mm (n=1), greyish brown; broad, ratio of length to breadth 3.2; R1 slightly curved before crossvein r; radial sector not produced anterad immediately before discoidal cell; discoidal cell originating at mid-length of wing, about 3x longer than wide; median cell slightly shorter than discoidal cell; crossvein r situated at base of fork I; fork I originating before crossvein s at distance equal to length of crossvein s; nygma located near base of fork II; fork III 1 /5th as long as wing; fork V as long as fork I, shorter than fork II; Cu2 nearly fused with A at wing margin. Hind wings 4.7 mm (n=1), grey; broad, ratio of length to breadth 2.9; margin weakly incurved at arculus, where Cu1 and Cu2 fused with margin; fork III slightly longer than discoidal cell and 1/7th as long as wing; fork V as long as fork I; 1A+2A 5x longer than 1A.

Male genitalia ( Figs. 155–159 ). Segment IX slightly shorter than high; longitudinally oval except posterior part expanded dorsally; each anterodorsal margin deeply concave in lateral view; each ventral margin convex, with strong incision at vertical apodeme; each posterior margin slightly convex, starting below cercus; ventral side of posterior 1/2 of segment IX with setae ( Fig. 157 ). In dorsal view segment IX with narrow anterior lobes; in dorsal view anterodorsal margin forming deep U-shaped incision with anterad-orienting processes on each side. In ventral view segment IX nearly quadrangular, with shallowly concave anterior margin and straight posterior margin; posterior margin without central projection. Tergum X simple; tapering apically from mid-length in lateral view ( Fig. 155 ); with dorsal process at mid-length; orienting posteroventrad, nearly straight; divided into 2 lateral branches from base in dorsal and ventral view (Figs. 156,157); each branch uniformly narrowing posteriorly in dorsal and ventral view, apically pointed, without processes; pair of sensillae located on dorsal part of each apex. Cerci large, cylindrical, located well below dorsal margin of segment IX and tergum X and oriented posterodorsad in lateral view; covered by long setae. Gonopods as long as segment IX, 3-branched. Dorsal branch of each gonopod long, exceeding tergum X in lateral view; dorsal branch uniformly slender, uniformly curved posteroventrad; anterodorsal margin convex; ventral margin slightly concave; in ventral view slightly converging before apex; apex running parallel, with small mesal megasetae. Ventral branch of each gonopod hyperbolic in lateral view, with undulating posterior margin and more smooth ventral margin; in ventral view, mesal margins nearly straight, strongly diverging. Mesal branches darkly pigmented, about 2x longer than each circus; in lateral view curved posterad before mid-length; in ventral view curved mesad at mid-length. Phallic apparatus slightly longer than rest of genitalia: phallotheca, in lateral view, with anterior part nearly 3x thicker than posterior part; and in ventral view anterior part about 2x wider than posterior part; apicoventral spine absent on phallotheca; small phallotremal sclerite in phallic apparatus forming U-like structure about as long as endothecal spines; 2 short, nearly black endothecal spines present.

Female. Unknown.

Holotype male: TAVEUNI: Cakadrove Prov., Devo Peak Radio Tower, Malaise trap in rain forest, 25.ix–7.x.2002, 16°51’S, 179°58’E [16.8500°S, 179.9667°E], 1200 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#36] [ FNIC].

Paratypes: Same data as holotype, except 13–20.xii.2002 [loc#36] — 4 males [ NHRS]. Cakaudrove Prov., Devo Forest Reserve, Malaise trap, 3–10.i.2003, 16°50’S, 179°59’E, 800 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#37] — 1 male [ BPBM]. Same data, except 10–16.i.2003 [loc#37] — 4 males [ BPBM]. Cakadrove Prov., 5.5 km SE Tavuki Village, Malaise trap in rainforest, 30.iv–14.viii. 2004,16.843°S, 179.955°W, 1188 m, E. Schlinger & M. Tokota’a [loc#38] — 1 male [ NHRS, DNA voucher IN1]. VANUA LEVU : Bua Prov., Kilaka, Malaise trap, 28.vi–2.vii.2004, 16°48’412’’S, 178°59’017’’E [16.8114°S, 178.9839°E], 412 m, M. Irwin, E. Schlinger & M. Tokota’a [loc#23] — 3 males [ NHRS].

Etymology: Psychodida , named after the Diptera family Psychodidae (moth flies) into which the genus Trichoptera Meigen was described.

Distribution: Taveuni and Vanua Levu.

Figures

FIGURES 22 – 29. Right wings, dorsal views. 22 — C. cartwrighti, new species, holotype; 23 — C. kimminsi, new species, holotype; 24 — C. kadavuensis, new species, holotype; 25 — C. lavensis, new species, holotype; 26 — C. devoensis, new species, holotype; 27 — C. helomyzida, new species, holotype; 28 — C. tipulida, new species, holotype; 29 — C. psychodida, new species, holotype.

FIGURES 155 – 159. Chimarra psychodida, new species, holotype. 155 — genitalia, left lateral view; 156 — genitalia, dorsal view; 157 — genitalia, ventral view; 158 — phallic apparatus, left lateral view; 159 — phallic apparatus, ventral view.