Ptomaphaminus testaceus Schilthuizen & Perreau

Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva, 2018, A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae), ZooKeys 777, pp. 57-108 : 81-82

Ptomaphaminus testaceus Schilthuizen & Perreau sp. n. Figures 11e, 12c, d, 13c, d

Material.

Holotype: Malaysia, Sarawak, 4th Division, Gunung Mulu National Park, mixed dipterocarp forest, v–viii.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), male (aedeagus on separate microscope slide). Paratypes: Sarawak. Gunung Mulu National Park, pitfall-trap, fish bait, alluv. for., ca. 100 m, iii–v.1978 (leg. I. Hanski, NHMUK, B.M. 1978-524), 2 males, 4 females; Gunung Mulu National Park, mixed dipterocarp forest, v–viii.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), 1 female; Gunung Mulu National Park, 4.5.78, Tps 7-13, v–viii.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), 1 female.

Description.

Length: 1.4-1.8 mm. General colour light reddish brown; legs, antenna, and mouthparts yellowish. Winged. Eyes well developed. Pronotum 1.45 times as wide as long. Elytra 1.45 times as long as wide. Female sutural angle of elytra expanded backwards into a sharp tooth. Male protarsi 0.4 times as wide as the apex of protibia. Male genital segment with a very elongate thin spiculum gastrale (Figure 13c). Aedeagus approximately 3.25 times shorter than the body length, elongate and parallel-sided, apex rounded, with two overlapping apical right and left expansions in dorsal view (Figure 12d), regularly arcuate from base to apex, without apical hook in lateral view (Figure 12c). Two lateral setae on each side, orthogonal to the plane of the aedeagus. Internal stylus of the endophallus short and strongly sinuate, confined to the apical half of the length of the median lobe (Figure 12c). Female genital segment with normally developed gonocoxites. Spermiduct slightly helical. Spermatheca weakly sclerified, with a set of transverse rings and without apical sclerotised plate (Figure 13d).

Differential diagnosis.

Very similar to P. latescens , but distinct in the external morphology by its significantly smaller size, and its lighter colour. The spermiduct is slightly helical, which is not the case in P. latescens .

Habitat and distribution.

Known only from the lowland forest of Gunung Mulu, Sarawak, Malaysia.

Etymology.

Named for its light brown colour ( testaceus = brick-coloured).