Cratotabanus Martins-Neto and Santos

do Carmo, Daniel Dias Dornelas, Sampronha, Stephanie, Santos, Charles Morphy D. & Ribeiro, Guilherme Cunha, 2022, Cretaceous Horse flies and their phylogenetic significance (Diptera: Tabanidae), Arthropod Systematics & amp; Phylogeny 80, pp. 295-307 : 295

Cratotabanus Martins-Neto and Santos

Figs 1 , 2 , 3 , 4

Cratotabanus Martins-Neto and Santos 1994: 291, fig. 1, pl. 1; Grimaldi et al. 2011: 303 (revised diagnosis); Grimaldi 2016: 45, fig. 16A (emended diagnosis).

Diagnosis.

Ocellar triangle developed with visible ocelli; basal callus absent; postspiracular scale absent; R5 slightly curved; hind tibia with 2 spurs; tergite X undivided; cercus two-segmented.

Comments.

The genus Cratotabanus differs from extant horse flies by the presence of two-segmented cercus and by the absence of the postspiracular scale, a one-segmented cercus and a postspiracular scale is shared by all extant Tabanidae ( Mackerras 1954). Grimaldi (2016) also pointed out that the length of the discal cell in Cratotabanus is nearly the same as the length of the vein M1. However, this character is variable in the extant species. The genus can be attributed to Tabanidae by the general disposition of the wing veins, the general shape of the body, the clypeus format and the presence of an enlarged calypter ( Grimaldi 2016, and our Figs 1 , 2 ).

Cratotabanus differs from Laiyangitabanus Zhang and Eotabanoid Mostovski et al. by the presence of vein R2+3 curved only at the apex. From Laiyangitabanus , Cratotabanus further differs by the presence of a well-developed ocellar triangle, and r-m closer to the base of the discal cell ( Grimaldi 2016). Cratotabanus differs from Eopangonius Ren by the absence of basal callus and a not projected clypeus.

Figures

Figure 1. Cratotabanus stonemyomorphus. Martins-Neto and Santos. A, B Holotype female. A habitus preserved in ventral view. B Detail of head. C Male, preserved in dorsal view. Scale bars: 2 mm (A, C); 1 mm (B).

Figure 2. Cratotabanus stonemyomorphus. A Holotype female, illustration of dorsal habitus. B Male, illustration of ventral habitus. C Holotype female, wing. D Male, Wing. Scale bars: 2 mm (A, C); 1 mm (C, D).

Figure 3. Cratotabanus cearensis sp. n. Holotype female. A Lateral habitus. B Head and thorax. C Terminalia. Scale bars: 2 mm (A); 1 mm (B); 0.5 mm (C)

Figure 4. Cratotabanus cearensis sp. nov. Holotype female. A Illustration of lateral habitus. B Wing venation. C Female terminalia. Scale bars: 2 mm (A, B); 0.1 mm (C).