Paraphloeostiba coriacea ( Cameron, 1952 )

Shavrin, Alexey V., 2024, Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960, Zootaxa 5424 (3), pp. 251-307 : 271-272

Paraphloeostiba coriacea ( Cameron, 1952)

( Figs 39 , 45 , 53–56 )

Phloeonomus coriaceus Cameron, 1952: 243

Paraphloeostiba coriacea : Steel 1960: 146, Shavrin 2023b: 489

Material examined. INDONESIA: WESTERN NEW GUINEA: 55 ♂♂, 53 ♀♀: ‘ IRIAN JAYA : Paniai Prov. Bilogay, 2100-2200 m 29.12.1995 leg. A. Riedel’ ( NMW, cSh); 2 ♂♂, 3 ♀♀: ‘ IRIAN JAYA : Paniai Prov. Bilogay, Tigipigu 2100-2400m, 23. 12. 1995 leg. A. Riedel’ ( NMW); 8 ♂♂, 11 ♀♀: ‘ IRIAN JAYA : Paniai Prov. Sinak, 2000-2200m 14.- 17.12.1995 leg. A. Riedel’ (2 ♂♂, 2 ♀♀: cSh; 6 ♂♂, 8 ♀♀: NMW); 1 ♀: ‘ IRIAN JAYA : Paniai Prov. | Sinak, trail to Ilaga | 2100-2500m, 15. 12. 1995 | leg. A. Riedel’ <printed> ( NMW); 1 ♀: ‘ NEW GUINEA Indona [Sefuna] 17.XI.1965 ’, ‘ R . Hornabrook | BM 1970-232 .’ ( BMNH) .

Redescription. Measurements (n=133, including the holotype ( Shavrin 2023b)): HW: 0.43–0.52; HL: 0.24– 0.36; OL: 0.14–0.17; TL: 0.02–0.06; AL (holotype): 0.85; PL: 0.32–0.42; PWmax: 0.52–0.69; PWmin: 0.47–0.66; ESL: 0.63–0.76; EW: 0.76–0.91; MTbL (holotype): 0.34; MTrL (holotype): 0.16 (MTrL 1–4: 0.05; MTrL 5: 0.11); AW: 0.69–0.87; AedL: 0.42–0.50; BL: 2.08–3.08 (holotype: 2.56).

Habitus as in Fig. 2 in Shavrin (2023b) and in Fig. 39 . Body brown, with paler lateroapical portion of head, lateral and basal portion of elytra (some specimens with uniformly paler elytra in contrast to the body), paratergites and apical tergites of abdomen in some specimens (some specimens with slightly darker abdominal tergites V–VII in middle); antennomeres 6–11 yellow-brown to brown; mouthparts, antennomeres 1–5 and legs yellowish (two preapical segments of maxillary palpomeres distintly darker in some specimens). Head (including neck), pronotum and scutellum without punctures; elytra with moderately fine and dense punctation, sometimes finer along suture and indistinct in apical portions; abdomen with very indistinct dense punctation. Head with regular and very strong coarse microsculpture, sometimes slightly finer in middle and coarser on infraorbital portions; neck with fine or relatively coarse and dense isodiametric or transverse meshes; microsculpture of pronotum about as that in middle portion of head or slightly stronger and coarser; scutellum with dense and coarse transverse or isodiametric meshes; microsculpture of elytra isodiametric and very dense, slightly finer and sometimes larger than that in pronotum, usually denser and finer in parascutellar portion and along suture; abdomen with very dense isodiametric microsculpture, distinctly transverse in abdominal tergites VI–VIII in some specimens.

Head 1.4–1.7 times as broad as long, with moderately wide and deep anteocellar foveae, sublinear or slightly diverging posteriad, about as long as diameter of ocellus or slightly longer, reaching level of posterior third or about middle length of eyes; postocular carina absent or very indistinct, obtuse. Ocelli moderately large, located about at level of postocular ridges; distance between ocelli about as long as or slightly greater than distance between ocellus and posterior margin of eyes. Apical maxillary palpomere about twice as long as preapical segment, or slightly shorter. Antennomere 4 distinctly longer than broad, 5 slightly broader than 4, 6 distinctly broader than 5, 7 slightly broader and longer than 6, 8–9 slightly shorter than 7, 10 slightly longer than 9, apical antennomere about 1.2 times as long as 10.

Pronotum 1.6–1.7 times as broad as long, 1.2–1.3 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; posterior angles subacute or obtuse; middle portion with slightly impressed medioapical portion in some specimens and with two indistinct to distinct wide and moderately long paramedian impressions.

Elytra 1.1–1.3 times as broad as long, 1.8 times to twice as long as pronotum, with narrowly impressed lateral margins.

Abdomen distinctly narrower than elytra.

Male. Posterior margin of abdominal tergite VIII truncate, as in Fig. 13 in Shavrin (2023b). Posterior margin of abdominal sternite VIII slightly concave, as in Fig. 14 in Shavrin (2023b). Aedeagus with wide basal bulb, strongly narrowed toward moderately small truncate apex; parameres distinctly exceeding apex of median lobe, gradually widened apically, with two long and two moderately short apical setae, and with a row of 7–8 of moderately short setae on inner margins and one short seta on outer margins of each paramere; internal sac narrow and moderately short, with several sclerotized elongate stuctures in basal portion (see Fig. 11 in Shavrin 2023b and Fig. 53 ). Lateral aspect of the aedeagus as in Fig. 12 in Shavrin (2023b).

Female. Posterior margins of abdominal tergite VIII ( Fig. 54 ) and abdominal sternite VIII ( Fig. 55 ) truncate. Female accessory sclerite as in Fig. 56 . Spermatheca not recognized.

Comparative notes. See details in Shavrin (2023b).

Distribution. Paraphloeostiba coriacea is known from several localities in Western New Guinea ( Indonesia) and Papua New Guinea.

Bionomics. The species was collected at elevations from 1500 (holotype) to 2500 m a.s.l. The detailed bionomical data are unknown.

Remarks. Based on the study of the holotype, this species was redescribed by Shavrin (2023b). I provided here the redescription this species, including the description of the female. It is here recorded from the Western New Guinea ( Indonesia) for the first time.

Figures

FIGURE 11. Distribution of Nipponophloeostiba: N. verrucifera (circles), N. thayeri (triangle), N. longilobata (square), N. explanata (diamond), N. hornabrooki (star).

FIGURES 45. Distribution of Paraphloeostiba coriacea (circles) and P. gayndahensis (squares) in New Guinea.

FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm.

FIGURES 53–56. Paraphloeostiba coriacea: 53–aedeagus, parameral view (Western New Guinea), 54–female abdominal tergite VIII, 55–female abdominal sternite VIII, 56–female accessory sclerite. Scale bar: 0.1 mm.

FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm.

FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm.