Cacodaemon laotinus laotinus (Arrow, 1920)

Yoshitomi, Hiroyuki & Pham, Thai Hong, 2022, The genus Cacodaemon (Coleoptera, Endomychidae) of Vietnam, ZooKeys 1081, pp. 127-136 : 127

publication ID	https://dx.doi.org/10.3897/zookeys.1081.75927
publication LSID	lsid:zoobank.org:pub:AD2D54F5-56D1-4DB2-B7F8-F3B3E0528394
persistent identifier	https://treatment.plazi.org/id/A2375B97-C35B-5682-8B59-63F249F49EB7
treatment provided by	ZooKeys by Pensoft
scientific name	Cacodaemon laotinus laotinus (Arrow, 1920)
status

Treatment

Figs 1A-1D View Figure 1 , 2A-2D View Figure 2 , 3A-3D View Figure 3 , 4A-4H View Figure 4 , 5A View Figure 5 , 5B View Figure 5

Amphisternus bellicosus var. laotinus Arrow, 1920: 322.

Amphisternus laotinus : Arrow 1928: 343; Strohecker 1953: 110.

Cacodaemon laotinus : Strohecker 1964: 350; Shockley et al. 2009: 36.

Material examined.

1 male & 1 female (EUMJ), "[ Laos] East Nong Het Xieng Khouang Prov. 25. VI. 2006 J. Yamasako leg." ; 3 males & 1 female (NIAS, VNMN), " Tam Dao N. Vietnam June 1996 ", " T. Kumasawa Collection " ; 1 female (NIAS), " Tam Dao N. Vietnam 1 May 1997 ", " T. Kumasawa Collection " ; 4 males & 1 female (SEHU), " Tam Dao VI 1994 ", "A. TANAKA Coll. (田中明) Sehu Japan 2005" .

Diagnosis.

This is a distinct species in the genus by having the following characteristics: elytral appendages, apical parts of femur, and antennomere 1 tinged with orange/dull orange; DA conoidal, not simply pointed.

Redescription.

Male. Body (Fig. 1A View Figure 1 , 1C View Figure 1 ) oval, convex dorsally, weakly shiny. Coloration of body black; BA, DA, PA, humeral parts, and apical parts of femur orange; antennomere 1 I and anterolateral corners of pronotum dull orange.

Head finely punctate. Antennae long and slender; antennomere 3 longest, a little shorter than 4 and 5 combined; club (antennomeres 9-11) distinctly wide. Pronotum impunctate, microreticulate, widest at middle; basal and lateral grooves shallow; front corners projecting and pointed; lateral margins slightly tapered anteriorly and posteriorly; posterior corners slightly projecting posterolaterally; PLM/PLS 0.74-0.77 (0.75); PWM/PWA 1.14-1.23 (1.20); PWM/PLM 1.43-1.52 (1.49); PWM/PLS 1.06-1.17 (1.12). Fore tibia (Fig. 2A View Figure 2 , 2C View Figure 2 ) straight and long, slender, with a sharp denticle at middle of inner margin. Elytra (Fig. 3A View Figure 3 , 3C View Figure 3 ) finely and sparsely punctate, microreticulate; BA large tubercle; HA long spine, projecting laterally, stout in basal part; DA projecting dorsally, conoidal, with small spine or dull apex, stout in basal part; PA a pair of tubercles; apex of elytra with long spines; EL/EWH 0.90-0.92 (0.91); EL/PLM 3.43-3.51 (3.46); EWH/PWM 2.46-2.67 (2.58); EWH/PWM 1.73-1.67 (1.71); TL/EWH 1.15-1.19 (1.17). Aedeagus (Fig. 4 View Figure 4 ) stout; apical branch (ab) short, with acute apex; subapical branch (sb) long, slightly curved, with pointed apex.

Female (Figs 1B View Figure 1 , 1I View Figure 1 , 3B View Figure 3 , 3D View Figure 3 ). Sexual dimorphism distinct in the following characteristics: fore tibia (Fig. 2B View Figure 2 , 2D View Figure 2 ) straight and long, lacking denticle; posterior corners of pronotum right-angled. PLM/PLS 0.73-0.74 (0.74); PWM/PWA 1.26-1.27 (1.27); PWM/PLM 1.52-1.57 (1.54); PWM/PLS 1.13-1.14 (1.14); EL/EWH 0.92; EL/PLM 3.43-3.74 (3.59); EWH/PWM 2.46-2.58 (2.52); EWH/EWM 1.76-1.72 (1.74); TL/EWH 1.17-1.19 (1.18). Sternite VIII (Fig. 5A View Figure 5 ) deeply emarginate at posterior margin. Ovipositor (Fig. 5B View Figure 5 ) bearing long setae along lateral margins of fused coxites; posterior margin of fused coxites arcuate; styli bearing long setae.

Measurement. Male from Vietnam (N = 3). TL 10.20-10.45 (10.34) mm; PWM 3.30-3.58 (3.44) mm; PWA 2.80-2.90 (2.87) mm; PLM 2.30-2.35 (2.32) mm; PLS 3.05-3.10 (3.08) mm; EL 7.90-8.10 (8.03) mm; EWH 8.80-9.00 (8.87) mm; EWM 5.10-5.40 (5.20) mm. Male from Laos (N = 1). TL 10.10 mm; PWM 3.45 mm; PWA 2.80 mm; PLM 2.30 mm; PLS 3.10 mm; EL 7.80 mm; EWH 8.28 mm; EWM 5.00 mm. Female from Vietnam (N = 2). TL 10.20-10.90 (10.55) mm; PWM 3.50-3.60 (3.55) mm; PWA 2.75-2.85 (2.80) mm; PLM 2.30 mm; PLS 3.10-3.15 (3.13) mm; EL 7.90-8.60 (8.25) mm; EWH 8.60-9.30 (8.95) mm; EWM 4.90-5.40 (5.15) mm. Female from Laos (N = 1). TL 9.00 mm; PWM 3.25 mm; PWA 2.70 mm; PLM 2.00 mm; PLS 3.00 mm; EL 7.00 mm; EWH 7.60 mm; EWM 4.70 mm.

Distribution.

Laos, Vietnam (new record).

Remarks.

As already mentioned by Strohecker (1964), Cacodaemon laotinus yunnanensis (Kryzhanovskij, 1960) described from Yunnan (China), is thought to be an infraspecific variation. We do not treat this subspecies in this paper because the holotype could not be examined.