Helonoton tico Lord and Ivie, 2016

Lord, Nathan P. & Ivie, Michael A., 2016, Several New Genera and Species of New World Synchitini (Coleoptera: Zopheridae: Colydiinae), The Coleopterists Bulletin 70 (4), pp. 715-753 : 715-753

Helonoton tico Lord and Ivie , new species ( Figs. 16 , 24 )

Diagnosis. Helonoton tico is most similar to H. foleyi in general appearance and possessing narrow, elongate carinae at the base and apex of the elytra, but can be distinguished by the absence of a tubercle/short carina on interstrial interval 7 at the elytral mid-point, the central carinae on pronotal disc strongly raised, the elevation of the central portion greater than the anterior and posterior carinae on the disc, and the Costa Rican distribution.

Description. Size moderate (TL = 4.0 mm), body elongate (TL/EW = 2.5), parallel-sided; light brown to dark brown; habitus as in Fig. 16 . Head: Elongate (HW = 0.7 mm), not constricted behind eyes. Antennae: 11-segmented, ending in an abrupt, 2-segmented club; antennomeres 10– 11 forming distinct club, antennomere 10 transverse, trapezoidal to slightly asymmetrical, narrowest at base; antennomere 11 truncate at base, transverse, rounded apically. Prothorax: Pronotum subquadrate to elongate (PW = 1.2 mm, PL = 1.1 mm, PL/ PW = 0.9), widest at apical ¼, narrowest at base. Lateral margins of pronotum sinuate, widening apically, anterior ⅓ produced into distinct arcuate lobe, lateral margin with constriction at middle, posterior ½ sinuate, microdenticulate; anterior angles distinct, angulate, projecting forward to slightly past level of anterior margin; posterior angles weak, angulate. Pronotal disc with complex patterns of ridges and depressed areas; central portion with depressed area, bordered laterally by sinuate ridge, ridge equally raised throughout; central portion bordered anteriorly by paired, short, parallel ridges that become confluent with anterior margin; basal portion of central discal area with paired, baso-laterally directed ridges that end before posterior margin, creating a subtriangular baso-medial depression. Pair of mid-lateral, weakly angulate tubercles present between lateral pronotal margin and basal ½ of central sinuate tuberculate ridge, as well as smaller pair of angulate tubercles present between lateral pronotal margin and anterior portion of central sinuate tuberculate ridge; posterior pair slightly larger than anterior pair. Procoxal cavities broadly open externally. Mesothorax: Scutellum well-developed, visible, ovoid to pentagonal; anterior margin rounded to truncate, posterior margin rounded to slightly angulate. Elytra: Elongate, parallel-sided, widest at middle (EL = 2.9 mm, EW = 1.6 mm, EL/EW = 1.9). Surface with elongate tubercles or short carinae. Sutural stria raised, beaded, abruptly diverging antero-laterally to form scutellary striole. Each elytral base with 3 elongate carinae; 1 st at base of elytral interstrial interval 3, raised, elongate, distinctly swollen posteriorly; 2 nd at base of interval 5, ½ as long as carina on interval 3; 3 rd at base of interval 7, beginning at humeral angle and extending towards apex, 2X as long as carina on interval 5. Interval 7 in middle of elytra weakly raised, without distinct tubercle. Interval 9 without distinctly raised carina. Remaining tubercles on middle portion of elytra small and round to elongate-oval, subequal in size. Carinae present on elytral declivity, long, extending nearly to apex. Epipleuron present, weakly defined, incomplete to apex, ending at abdominal ventrite IV. Metaventrite: Longer than abdominal ventrite I, with paired sinuate grooves directly posteriad and bordering mesocoxae, also with paired, slightly curved grooves directly anteriad and bordering metacoxae. Discrimen moderate, extending to middle ½ of metaventrite, strongly impressed between and anteriad metacoxae. Metacoxae transverse, narrowly separated. Metendosternite of fully winged form as in Fig. 2g (see genus-group description). Abdomen: Intercoxal process of abdominal ventrite I triangular, apex acute. In males, ventrites III and IV with large, smooth, transverse-oval patches in lateral areas. Metathoracic wing: Fully developed.

Distribution. Costa Rica ( Fig. 24 ).

Biology. One specimen was collected in primary forest.

Etymology. Named in honor of Ticos, a colloquial term for the people of Costa Rica.

Type Material. 2 specimens. Holotype, female (pointed, KSEM): COSTA RICA: Heredia; Cerro

Chompipe ca. 2km. N.; Monte de la Cruz, 1950m; 10°5′13″N, 84°4′45″W; 25 JUL 2000; J.Ashe, R. Brooks,; Z.Falin CR1ABF00 241 / [label with barcode] SM0205773; KUNHM-ENT. Paratype, male (pointed, partially dissected, genitalia and abdomen in glycerine in genitalia vial pinned underneath specimen, TAMU): COSTA RICA: Prov. Heredia; 16km SSE La Virgen , 1050-; 1150m, 10°16′N 84°05′W; III-9-14-2001, E.G. Riley; primary forest .

Figures

Figs. 13–16. Helonoton species, dorsal habitus. 13) H. pascoei (MAIC); 14) H. pustulosum, holotype (FSCA);

Figs. 23–24. Distribution maps of Helonoton species. 23) Species in Mexico, Guatemala, El Salvador, and Honduras;

Fig. 2. Helonoton costaricense. a) Head, ventral view, b) Labium, ventral view, c) Maxilla, ventral view, d) Antenna, e) Fore leg, f) Mandible, ventral view, g) Metendosternite, ventral view, h) Prothorax, ventral view, i) Pterothorax, ventral view, j) Abdominal ventrites I–V, ventral view, k) Aedeagus, dorsal view, l) Aedeagus, lateral view.