Helonoton chiriqui Lord and Ivie, 2016

Lord, Nathan P. & Ivie, Michael A., 2016, Several New Genera and Species of New World Synchitini (Coleoptera: Zopheridae: Colydiinae), The Coleopterists Bulletin 70 (4), pp. 715-753 : 715-753

Helonoton chiriqui Lord and Ivie , new species ( Figs. 9 , 24 )

Diagnosis. Helonoton chiriqui most closely resembles H. pascoei , but can be identified by the non-interrupted scutellary striole, shorter, more ovoid elytral tubercles, less strongly raised central carina of the pronotal disc, and Panamanian distribution. Helonoton chiriqui also superficially resembles H. tatumbla , but the latter can be easily distinguished by the 3-segmented antennal club and head with distinct temples.

Description. Size moderate to large (TL = 3.5 mm), body elongate (TL/EW = 2.4), parallelsided; light brown to dark brown; habitus as in Fig. 9 . Head: Elongate (HW = 0.6 mm), not constricted behind eyes. Antennae: 11-segmented, ending in an abrupt, 2-segmented club; antennomeres 10–11 forming distinct club, antennomere 10 transverse, trapezoidal to slightly asymmetrical, narrowest at base; antennomere 11 truncate at base, transverse, rounded apically. Prothorax: Pronotum subquadrate to elongate (PW = 1.2 mm, PL = 1.0 mm, PL/PW = 0.9), widest at apical ¼, narrowest at base. Lateral margins of pronotum sinuate, widening apically, anterior ⅓ produced into moderate arcuate lobe, lateral margin with constriction at middle, posterior ½ subtly sinuate, microdenticulate; anterior angles dis- tinct, rounded, slightly projecting forward to about level of anterior margin; posterior angles weak, angulate. Pronotal disc with complex patterns of ridges and depressed areas; central portion with depressed area, bordered laterally by pair of outwardly arcuate ridges, ridge equally raised throughout; central portion bordered anteriorly by paired, short, parallel ridges that become confluent with anterior margin; basal portion of central discal area with paired, baso-laterally directed ridges that end before posterior margin, creating subtriangular baso-medial depression. Pair of short, mid-lateral carinae present between lateral pronotal margin and basal half of central sinuate tuberculate ridge, as well as smaller pair of shorter carinae present between lateral pronotal margin and anterior portion of central sinuate tuberculate ridge. Procoxal cavities broadly open externally. Mesothorax: Scutellum well-developed, visible, quadrate to ovoid; anterior margin truncate, posterior margin rounded to straight. Elytra: Elongate, parallel-sided, widest at middle (EL = 2.5 mm, EW = 1.5 mm, EL/EW = 1.7). Surface with elongate tubercles or short carinae. Sutural stria raised, beaded, abruptly diverging antero-laterally to form scutellary striole. Each elytral base with 3 elongate carinae; 1 st at base of elytral interstrial interval 3, raised, elongate, distinctly swollen posteriorly; 2 nd at base of interval 5, ⅔ as long as carina on interval 3; 3 rd at base of interval 7, more weakly defined, beginning at humeral angle and extending towards apex, 2X as long as carina on interval 5. Interval 9 without distinctly raised carina. Remaining tubercles on middle portion of elytra small and round to elongate-oval, of variable sizes. Carinae present on elytral declivity, long, extending nearly to apex. Epipleuron present, weakly defined, incomplete to apex, ending near middle of abdominal ventrite IV. Metaventrite: Longer than abdominal ventrite I, with paired sinuate grooves directly posteriad and bordering mesocoxae, also with paired, slightly curved grooves directly anteriad and bordering metacoxae. Discrimen, extending to middle half of metaventrite, strongly impressed between and anteriad metacoxae. Metacoxae transverse, narrowly separated. Metendosternite of fully winged form as in Fig. 2g (see genus-group description). Abdomen: Intercoxal process of abdominal ventrite I triangular, apex acute. In males, ventrites III and IV with smooth, slightly raised, transverse-oval patches in lateral areas; patches less obvious than in other species. Metathoracic wing: fully developed.

Distribution. Panama ( Fig. 24 ).

Biology. One specimen was collected from a gilled mushroom.

Etymology. Named after the Chiriquí province in western Panama, the locality where the type series was collected.

(MNCR); 11) H. foleyi , holotype (CNCI); 12) H. mexicanum , paratype (MAIC). Scale bars = 1 mm.

Type Material. 10 specimens. Holotype, male (pointed, KSEM): PANAMA: Chiriqui Prov.; Volcan Baru ; 1 km NW Boquete; 08°48′N, 82°29′W; 2200 m, 18 VI 1995 ; J.Ashe&R. Brooks#241; ex: gilled mushrooms. Paratypes (3 pointed, CNCI): PANAMA, Chiriqui; Prov. 2-3km E Cerro; Punta , 2000-2200m , 23.; V.77 H. & A. Howden. Paratypes (3 pointed, one male partially dissected, genitalia and abdomen in glycerine in genitalia vial pinned underneath specimen, CNCI): PANAMA, Chiriqui; Prov. 2-3km E Cerro; Punta , 2000-2200m , 28.; V.77 H. &A. Howden. Paratypes (2 pointed, CNCI): PANAMA, Chiriqui; Prov. 2-3km E Cerro; Punta , 2000-2200m , 1.; VI.77 H. & A. Howden. Paratype (pointed, CNCI): PANAMA, Chiriqui; Prov. 2-3km E Cerro; Punta , 2000-2200m , 8.; VI.77 H. & A. Howden.

Figures

Figs. 9–12. Helonoton species, dorsal habitus. 9) H. chiriqui, holotype (KSEM); 10) H. costaricense, holotype

Figs. 23–24. Distribution maps of Helonoton species. 23) Species in Mexico, Guatemala, El Salvador, and Honduras;

Fig. 2. Helonoton costaricense. a) Head, ventral view, b) Labium, ventral view, c) Maxilla, ventral view, d) Antenna, e) Fore leg, f) Mandible, ventral view, g) Metendosternite, ventral view, h) Prothorax, ventral view, i) Pterothorax, ventral view, j) Abdominal ventrites I–V, ventral view, k) Aedeagus, dorsal view, l) Aedeagus, lateral view.