Ummidia gabrieli, Godwin & Bond, 2021

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia gabrieli sp. nov. Figs 34 , 35 , Map 4

Type material.

HOLOTYPE ♂ (UMM341) from 8 miles southeast of La Paz, La Paz, Baja California Sur, Mexico, 24.0345 -110.227 4, 304 m a.s.l., 13.x.1968, coll. EL Sleeper, FJ Moore, AMNH. PARATYPE: 1 ♀ (UMM392) from 5 miles north of San Ignacio, Mulegé, Baja California Sur, Mexico, 26.735 -111.7012 5, 210 m a.s.l., 20.i.1965, coll. V. Roth, AMNH.

Etymology.

The specific epithet is a patronym in honor of musician and human rights activist Peter Gabriel.

Diagnosis.

Ummidia gabrieli are relatively large spiders (males CL 8.6-10.6, other species CL 4-7.5) and can be differentiated from males of all other geographically proximal species by the lack of prolateral spines on metatarsus and tibia I, having a palpal tibia that is> 3 ' long as wide, by having tarsi which are relatively short and tapered distally, and by the presence of a dorsal opalescence on the abdomen. Males can be differentiated from U. mercedesburnsae , U. paulacushingae , U. waunekaae , and U. gertschi by having a sinuous embolus. Females can be differentiated from U. gertschi , U. modesta , and U. mercedesburnsae by having spermathecae that curve laterally. Females can be differentiated from U. pesiou by having significantly fewer endite cuspules (28 vs 59) and having rastellar spines not extending up cheliceral face, and from U. pustulosa by having the sternum roughly as wide as long.

Description of male holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Left palp and leg I removed, in vial with specimen. General coloration. Carapace, chelicerae, and legs reddish black 2.5YR 2.5/1. Abdomen black 2.5Y 2.5/1. Cephalothorax. Carapace 9.0 long, 8.39 wide. Pars cephalica 5.49 long. Foveal groove procurved, 0.61 long, 1.71 wide. All eyes on defined moderate tubercle. AER procurved. PER straight. Eye group 1.09 long, 1.75 wide, AME 0.47, PME 0.31, ALE 0.69, PLE 0.22. Sternum sparsely setose, STRl 5.09, STRw 4.41. Chelicerae with anterior tooth row comprising eight teeth, posterior margin with eight8 teeth. Palpal endites with eleven small, non-hastate cuspules over proximal 1/4 of endite face, lacking distal endite cuspules, ENDw 1.89, ENDl 3.35. Labium with four cuspules, LBw 1.77, LBl 1.39. Rastellum with many small, rounded teeth on process. Abdomen setose. Legs. F1 8.04; F1w 1.94; P1 3.59; Ti1 5.82; Mt1 3.92; Tr1 1.79; F3 6.11; F3w 2.32; P3 3.23; Ti3 3.53; Sd3 2.71; Mt3 3.79; Tr3 2.59; F4 7.87; F4w 2.22; P4 3.44; Ti4 5.06; Mt4 6; Tr4 2.67. Retrolateral face of tarsus IV lacking distinct comb or brush, but very setose ventrally. Leg I spination pattern: TSp 0, TSpv 0, TSrd 0, TSr 0, TSrv 11, MtSp 0, MtSr 7, TrSp 0, TrSr 7. Pedipalps. PTl 4.84, PTw 1.51, Bl 3.82. Embolus relatively long; sinuous.

Variation, males

(n = 7). CL 8.59-10.61, 9.24 ± 0.26; CW 7.84-9.92, 8.5 ± 0.25; Cap 5.28-6.41, 5.65 ± 0.14; ENDl 0.91-1.09, 0.98 ± 0.02; ENDw 1.65-1.88, 1.75 ± 0.04; STRl 4.41-5.97, 5.17 ± 0.17; STRw 4.27-5.08, 4.52 ± 0.1; LBl 1.15-1.6, 1.38 ± 0.05; LBw 1.46-4.77, 2.13 ± 0.44; F1 6.92-9.06, 7.86 ± 0.24; F1w 1.75-2.32, 2 ± 0.07; P1 3.21-4.41, 3.77 ± 0.14; Ti1 4.69-6.52, 5.7 ± 0.21; Mt1 3.31-4.57, 3.89 ± 0.14; Tr1 1.62-2.01, 1.82 ± 0.05; F3 5.2-6.84, 6.02 ± 0.19; F3w 2.18-2.74, 2.37 ± 0.07; P3 2.78-3.65, 3.18 ± 0.1; Ti3 3.36-4, 3.61 ± 0.08; Mt3 3.03-4.34, 3.7 ± 0.16; Tr3 2.23-2.96, 2.52 ± 0.09; F4 7.01-8.91, 7.77 ± 0.23; F4w 1.92-2.48, 2.16 ± 0.06; P4 3.1-3.82, 3.43 ± 0.08; Ti4 4.32-5.65, 5.07 ± 0.15; Mt4 4.74-6.23, 5.56 ± 0.19; Tr4 2.36-3.06, 2.63 ± 0.1; TSp 0-0, 0 ± 0; TSpv 0-3, 0.86 ± 0.46; TSr 0-0, 0 ± 0; TSrv 5-17, 8.71 ± 1.63; PTl 3.91-5.48, 4.83 ± 0.19; PTw 1.45-1.62, 1.52 ± 0.02; BL 3.6-4.31, 3.78 ± 0.09.

Description of female paratype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. Right palp removed at patella, in vial with specimen. General coloration. Carapace, chelicerae, and legs dark reddish brown 5YR 3/1. Abdomen black 7.5YR 2.5/1, spinnerets dark yellowish brown 10YR 4/6. Cephalothorax. Carapace 8.25 long, 7.45 wide. Pars cephalica 5.48 long. Foveal groove procurved, 0.47 long, 1.67 wide. Eye tubercle low. AER procurved. PER recurved. Eye group 0.95 long, 1.63 wide, AME 0.35, PME 0.26, ALE 0.49, PLE 0.31. Sternum sparsely setose, STRl 4.81, STRw 4.25. Chelicerae with anterior row comprising seven teeth, posterior margin with seven teeth. Palpal endites with 18 cuspules spread across proximal half of endite and ten reduced cuspules distally, ENDw 1.97, ENDl 3.08. Labium with eight cuspules, LBw 1.75, LBl 1.23. Rastellum with many strong spines along cheliceral margin and on rastellar process. Abdomen. Evenly setose with pale speckles and slight dorsal opalescence. Legs. F1 4.62; F1w 1.77; P1 2.92; Ti1 2.84; Mt1 1.97; Tr1 0.95; F3 4.45; F3w 2.35; P3 2.94; Ti3 2.49, Sd3 1.63; Mt3 1.7; Tr3 1.54; F4 5.68; F4w 2.26; P4 2.99; Ti4 2.94; Mt4 2.94; Tr4 1.66. Retrolateral face tarsus IV with defined comb over length of tarsus. Pedipalps. PF 4.35, PP 2.5, PTi 2.6, PTr 2.16. Spermathecae curved laterally with distal coil, bulbs facing laterally.

Variation, females

(n = 3). CL 5.4-8.3, 7.32 ± 0.96; CW 4.27-7.58, 6.43 ± 1.08; Cap 3.55-5.56, 4.86 ± 0.66; ENDl 0.65-1.11, 0.9 ± 0.14; ENDw 0.88-1.64, 1.38 ± 0.25; STRl 3.22-5.21, 4.41 ± 0.61; STRw 2.6-4.63, 3.83 ± 0.62; LBl 0.82-1.48, 1.18 ± 0.19; LBw 1.15-1.75, 1.55 ± 0.2; F1 2.94-5.32, 4.3 ± 0.71; F1w 1.18-1.82, 1.59 ± 0.21; P1 1.82-3.18, 2.64 ± 0.42; Ti1 1.78-3.31, 2.64 ± 0.45; Mt1 1.22-2.28, 1.82 ± 0.32; Tr1 0.84-1.3, 1.03 ± 0.14; F3 2.55-4.71, 3.9 ± 0.68; F3w 1.29-2.4, 2.01 ± 0.36; P3 1.66-2.94, 2.48 ± 0.41; Ti3 1.46-2.51, 2.15 ± 0.35; Mt3 1.1-2.01, 1.61 ± 0.27; Tr3 1.2-1.63, 1.46 ± 0.13; F4 3.31-5.68, 4.88 ± 0.79; F4w 1.47-2.26, 1.97 ± 0.25; P4 1.9-3.31, 2.73 ± 0.43; Ti4 1.84-3.14, 2.64 ± 0.4; Mt4 1.78-3.4, 2.71 ± 0.48; Tr4 1.14-1.66, 1.46 ± 0.16; PF 2.71-5, 4.02 ± 0.68; PP 1.62-2.89, 2.34 ± 0.38; PTi 1.81-3.43, 2.62 ± 0.47; PTr 1.31-2.8, 2.09 ± 0.43.

Material examined.

Mexico: Baja California: Sur: 22 mi NE of Santo Domingo , 28.5223 -113.77246, 786 m a.s.l. (UMM0155, 15.ii.1966, 1♂, V Roth, AMNH) ; 8 mi SE La Paz, 24.0345 -110.2274, 304 m a.s.l. (UMM0340, 13.x.1968, 1♂, EL Sleeper, FJ Moore, AMNH); (UMM0341, 1♂, AMNH); (UMM0596, 1♂, AMNH); (UMM0637, 1♂, AMNH); (UMM0400, 1♂, AMNH); El Triunfo watercourse, 23.8099 -110.12411, 457 m a.s.l. (MY02696, 16.iii.2004, 1♀, M Hedin, BME); Hwy 286 (to San Juan de los Planes), 24.1003 -110.26921, 110 m a.s.l. (MY02695, 17.iii.2004, 1♀, M Hedin, BME); near La Paz , 24.1263 -110.29357, 11 m a.s.l. (UMM0687, vii.1990, 1♂, T Jackson, AMNH) ; 3 mi N Santa Anita , 23.2612 -109.71556, 110 m a.s.l. (UMM0445, 1.vii.1968, 1♀, AMNH) ; 5 mi N San Ignacio (mission), 26.735 -111.70125, 210 m a.s.l. (UMM0392, 20.i.1965, 1♀, V Roth, AMNH) .

Figures

Map 4. Locality records for western United States and northern Mexico species.

Figure 34. Ummidia gabrieli sp. nov. from Baja California Sur, Mexico. A male habitus illustration UMM 341 B female habitus illustration UMM 392. Scale bars: 4.0 mm.

Figure 35. Ummidia gabrieli sp. nov. from Baja California Sur, Mexico A-D male holotype (UMM 341) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female paratype (UMM 392). Scale bars: 1.0 mm.